Title	Instructors	Location	Time	Description	Cross listings	Fulfills	Registration notes	Syllabus	Syllabus URL
BIOL 010-301	Ecological Consequences of Climate Change	Arthur E. Dunham	LLAB 109	MW 02:00 PM-03:30 PM	Students will read and discuss seminal papers on a number of major topics in the ecology of climate change and the long-term consequences of the effects of climate change on ecological systems. Some of the topics include: effects of climate change on species distributions, disruption of plant pollinator systems and the consequences for ecosystem composition and stability, changes in the distribution and epidemiology of insect-borne infectious diseases, and the consequences of sea level rise and the increased intensity and frequency of severe weather events. Other topics may be covered. Grading will be based on participation in discussions, a paper on an approved topic, and a presentation on the topic of the student's paper.			Freshman Seminar
BIOL 017-001	The Biology of Food	R. Scott Poethig	LEVN AUD	MWF 01:00 PM-02:00 PM	This course will examine the ways in which humans manipulate - and have been manipulated by - the organisms we depend on for food, with particular emphasis on the biological factors that influence this interaction. The first part of the course will cover the biology, genetics, evolution, and breeding of cultivated plants and animals; the second part will concern the ecological, economic, and political factors that influence food production.		Living World Sector	Registration also required for Recitation (see below)
BIOL 017-201	The Biology of Food	R. Scott Poethig	LEVN AUD	F 02:00 PM-03:00 PM	This course will examine the ways in which humans manipulate - and have been manipulated by - the organisms we depend on for food, with particular emphasis on the biological factors that influence this interaction. The first part of the course will cover the biology, genetics, evolution, and breeding of cultivated plants and animals; the second part will concern the ecological, economic, and political factors that influence food production.			Registration also required for Lecture (see below)
BIOL 101-001	Intro Biology A	J. Nicholas Betley Doris Wagner	LEVN 111	MW 09:00 AM-11:00 AM	General principles of biology focusing on the basic chemistry of life, cell biology, molecular biology, and genetics in all types of living organisms. Particular emphasis will be given to links between the fundamental processes covered and current challenges of humankind in the areas of energy, food, and health.		Living World Sector	Registration also required for Laboratory (see below)
BIOL 101-101	Introduction To Biology A	Rong Guo	LLAB 114	M 02:00 PM-05:00 PM	General principles of biology focusing on the basic chemistry of life, cell biology, molecular biology, and genetics in all types of living organisms. Particular emphasis will be given to links between the fundamental processes covered and current challenges of humankind in the areas of energy, food, and health.			Registration also required for Lecture (see below)
BIOL 101-102	Introduction To Biology A	Rong Guo	LLAB 114	T 09:00 AM-12:00 PM	General principles of biology focusing on the basic chemistry of life, cell biology, molecular biology, and genetics in all types of living organisms. Particular emphasis will be given to links between the fundamental processes covered and current challenges of humankind in the areas of energy, food, and health.			Registration also required for Lecture (see below)
BIOL 101-103	Introduction To Biology A	Staver Bezhani	LLAB 114	T 01:00 PM-04:00 PM	General principles of biology focusing on the basic chemistry of life, cell biology, molecular biology, and genetics in all types of living organisms. Particular emphasis will be given to links between the fundamental processes covered and current challenges of humankind in the areas of energy, food, and health.			Registration also required for Lecture (see below)
BIOL 101-104	Introduction To Biology A	Staver Bezhani	LLAB 114	W 02:00 PM-05:00 PM	General principles of biology focusing on the basic chemistry of life, cell biology, molecular biology, and genetics in all types of living organisms. Particular emphasis will be given to links between the fundamental processes covered and current challenges of humankind in the areas of energy, food, and health.			Registration also required for Lecture (see below)
BIOL 102-001	Intro Biology B	Chris Jang Paul D. Sniegowski	LLAB 10	MWF 01:00 PM-02:00 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.		Living World Sector	Registration also required for Laboratory (see below)
BIOL 102-101	Introduction To Biology B	Staver Bezhani	LEVN L11	M 02:00 PM-05:00 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.			Registration also required for Lecture (see below)
BIOL 102-102	Introduction To Biology B	Lori Hynes Spindler	LEVN L12	M 02:00 PM-05:00 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.			Registration also required for Lecture (see below)
BIOL 102-103	Introduction To Biology B	H Luke Mckenna Anderson	LEVN L11	T 09:00 AM-12:00 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.			Registration also required for Lecture (see below)
BIOL 102-104	Introduction To Biology B	Lori Hynes Spindler	LEVN L12	T 09:00 AM-12:00 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.			Registration also required for Lecture (see below)
BIOL 102-105	Introduction To Biology B	H Luke Mckenna Anderson	LEVN L11	T 01:00 PM-04:00 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.			Registration also required for Lecture (see below)
BIOL 102-106	Introduction To Biology B	Lori Hynes Spindler	LEVN L12	T 01:00 PM-04:00 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.			Registration also required for Lecture (see below)
BIOL 102-107	Introduction To Biology B	Jessica A Ardis	LEVN L11	W 09:00 AM-12:00 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.			Registration also required for Lecture (see below)
BIOL 102-108	Introduction To Biology B	Staver Bezhani	LEVN L12	W 09:00 AM-12:00 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.			Registration also required for Lecture (see below)
BIOL 102-109	Introduction To Biology B	Jessica A Ardis	LEVN L11	W 02:00 PM-05:00 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.			Registration also required for Lecture (see below)
BIOL 102-110	Introduction To Biology B	Lori Hynes Spindler	LEVN L12	W 02:00 PM-05:00 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.			Registration also required for Lecture (see below)
BIOL 102-111	Introduction To Biology B	Staver Bezhani	LEVN L11	R 01:00 PM-04:00 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.			Registration also required for Lecture (see below)
BIOL 102-112	Introduction To Biology B	Karl G Siegert		CANCELED	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.			Registration also required for Lecture (see below)
BIOL 102-113	Introduction To Biology B	Staver Bezhani	LEVN L11	R 04:30 PM-07:30 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.			Registration also required for Lecture (see below)
BIOL 102-601	Intro Biology B	John E. Zimmerman	LLAB 109	F 05:00 PM-07:30 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.		Living World Sector	Course Must Be Taken For A Grade Registration also required for Laboratory (see below)		https://pennintouchdaemon.apps.upenn.edu/pennInTouchProdDaemon/jsp/fast.do?webService=syllabus&term=2019A&course=BIOL102601
BIOL 102-602	Introduction To Biology B	Jessica A Ardis	LLAB 101	F 02:00 PM-05:00 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.			Registration also required for Lecture (see below)
BIOL 102-603	Introduction To Biology B	Jessica A Ardis	LLAB 101	S 09:00 AM-12:00 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.			Registration also required for Lecture (see below)
BIOL 109-401	Intro To Brain & Behav	Julie Mcgurk	LEVN AUD	TR 01:30 PM-03:00 PM	Introduction to the structure and function of the vertebrate nervous system. We begin with the cellular basis of neuronal activities, then discuss the physiological basis of motor control, sensory systems, motivated behaviors, and higher mental processes. This course is intended for students interested in the neurobiology of behavior, ranging from animal behaviors to clinical disorders.	BIBB109401, PSYC109401	Living World Sector	Registration also required for Laboratory (see below)		https://pennintouchdaemon.apps.upenn.edu/pennInTouchProdDaemon/jsp/fast.do?webService=syllabus&term=2019A&course=BIOL109401
BIOL 109-402	Introduction To Brain and Behavior	Michelle Lynne Klima	LLAB 104	T 09:00 AM-10:30 AM	Introduction to the structure and function of the vertebrate nervous system. We begin with the cellular basis of neuronal activities, then discuss the physiological basis of motor control, sensory systems, motivated behaviors, and higher mental processes. This course is intended for students interested in the neurobiology of behavior, ranging from animal behaviors to clinical disorders.	BIBB109402, PSYC109402		Registration also required for Lecture (see below)
BIOL 109-403	Introduction To Brain and Behavior	Jennifer Ann Stiso	LLAB 104	T 10:30 AM-12:00 PM	Introduction to the structure and function of the vertebrate nervous system. We begin with the cellular basis of neuronal activities, then discuss the physiological basis of motor control, sensory systems, motivated behaviors, and higher mental processes. This course is intended for students interested in the neurobiology of behavior, ranging from animal behaviors to clinical disorders.	BIBB109403, PSYC109403		Registration also required for Lecture (see below)
BIOL 109-404	Introduction To Brain and Behavior	David Nathan White	LLAB 104	T 12:00 PM-01:30 PM	Introduction to the structure and function of the vertebrate nervous system. We begin with the cellular basis of neuronal activities, then discuss the physiological basis of motor control, sensory systems, motivated behaviors, and higher mental processes. This course is intended for students interested in the neurobiology of behavior, ranging from animal behaviors to clinical disorders.	BIBB109404, PSYC109404		Registration also required for Lecture (see below)
BIOL 109-405	Introduction To Brain and Behavior	Kevin Mitchell Goff	LLAB 104	T 03:00 PM-04:30 PM	Introduction to the structure and function of the vertebrate nervous system. We begin with the cellular basis of neuronal activities, then discuss the physiological basis of motor control, sensory systems, motivated behaviors, and higher mental processes. This course is intended for students interested in the neurobiology of behavior, ranging from animal behaviors to clinical disorders.	BIBB109405, PSYC109405		Registration also required for Lecture (see below)
BIOL 109-406	Introduction To Brain and Behavior	Aaron L Williams	LLAB 104	T 04:30 PM-06:00 PM	Introduction to the structure and function of the vertebrate nervous system. We begin with the cellular basis of neuronal activities, then discuss the physiological basis of motor control, sensory systems, motivated behaviors, and higher mental processes. This course is intended for students interested in the neurobiology of behavior, ranging from animal behaviors to clinical disorders.	BIBB109406, PSYC109406		Registration also required for Lecture (see below)
BIOL 109-407	Introduction To Brain and Behavior	Jared Perry Zimmerman	LLAB 104	F 10:30 AM-12:00 PM	Introduction to the structure and function of the vertebrate nervous system. We begin with the cellular basis of neuronal activities, then discuss the physiological basis of motor control, sensory systems, motivated behaviors, and higher mental processes. This course is intended for students interested in the neurobiology of behavior, ranging from animal behaviors to clinical disorders.	BIBB109407, PSYC109407		Registration also required for Lecture (see below)
BIOL 109-408	Introduction To Brain and Behavior	Sydney Cason	LLAB 104	F 12:00 PM-01:30 PM	Introduction to the structure and function of the vertebrate nervous system. We begin with the cellular basis of neuronal activities, then discuss the physiological basis of motor control, sensory systems, motivated behaviors, and higher mental processes. This course is intended for students interested in the neurobiology of behavior, ranging from animal behaviors to clinical disorders.	BIBB109408, PSYC109408		Registration also required for Lecture (see below)
BIOL 109-409	Introduction To Brain and Behavior	Daniel Richard Schonhaut	LLAB 104	F 03:00 PM-04:30 PM	Introduction to the structure and function of the vertebrate nervous system. We begin with the cellular basis of neuronal activities, then discuss the physiological basis of motor control, sensory systems, motivated behaviors, and higher mental processes. This course is intended for students interested in the neurobiology of behavior, ranging from animal behaviors to clinical disorders.	BIBB109409, PSYC109409		Registration also required for Lecture (see below)
BIOL 124-001	Intro Organism Biol. Lab	Linda J. Robinson	LEVN AUD	F 09:00 AM-10:00 AM	An intensive introductory laboratory course in organismal biology.			Registration also required for Laboratory (see below)
BIOL 124-101	Introductory Organismal Biology Lab	Linyang Ju	LLAB 101	M 02:00 PM-05:00 PM	An intensive introductory laboratory course in organismal biology.			Registration also required for Lecture (see below)
BIOL 124-102	Introductory Organismal Biology Lab	Chris Jang	LLAB 101	T 09:00 AM-12:00 PM	An intensive introductory laboratory course in organismal biology.			Registration also required for Lecture (see below)
BIOL 124-103	Introductory Organismal Biology Lab	Chris Jang	LLAB 101	T 01:00 PM-04:00 PM	An intensive introductory laboratory course in organismal biology.			Registration also required for Lecture (see below)
BIOL 124-104	Introductory Organismal Biology Lab	Yuqi Zhou	LLAB 101	W 09:00 AM-12:00 PM	An intensive introductory laboratory course in organismal biology.			Registration also required for Lecture (see below)
BIOL 124-105	Introductory Organismal Biology Lab	Linyang Ju	LLAB 101	W 02:00 PM-05:00 PM	An intensive introductory laboratory course in organismal biology.			Registration also required for Lecture (see below)
BIOL 124-106	Introductory Organismal Biology Lab	Yuqi Zhou	LLAB 101	R 01:00 PM-04:00 PM	An intensive introductory laboratory course in organismal biology.			Registration also required for Lecture (see below)
BIOL 203-601	Essentials of Biochem	Megan Ruth Elliott	FAGN 110	W 05:00 PM-08:00 PM	Intermediate level course covering principles of modern biochemistry. Topics include protein structure, protein purification and characterization, proteomics, enzyme kinetics and mechanisms, membrane structure and function, metabolism, and cellular energy transduction. Emphasis will be on biochemical problem solving, experimental design, and application of quantitative methods in a biological and clinical context. This course is open to students in the College of Liberal and Professional Studies only.			Course Must Be Taken For A Grade Registration also required for Recitation (see below)
BIOL 203-602	Essentials of Biochemistry	Megan Ruth Elliott	FAGN 110	W 08:00 PM-08:30 PM	Intermediate level course covering principles of modern biochemistry. Topics include protein structure, protein purification and characterization, proteomics, enzyme kinetics and mechanisms, membrane structure and function, metabolism, and cellular energy transduction. Emphasis will be on biochemical problem solving, experimental design, and application of quantitative methods in a biological and clinical context. This course is open to students in the College of Liberal and Professional Studies only.			Registration also required for Lecture (see below)
BIOL 204-001	Biochemistry	Philip Arthur Rea	STIT B26	MW 02:00 PM-03:30 PM	BIOL 204 examines the basic principles of protein structure, protein purification and characterization, proteomics, enzyme kinetics and mechanism, membrane structure and function, metabolism, and cellular energy transduction. The primary objective is to provide life scientists with an appreciation of basic principles of modern biochemistry, and of how the current conceptual and technical framework arose. Emphasis is placed on the experimental approaches and reasoning behind the dissection and reconstitution of these processes in a biological and, in some cases, clinical context. Discussions directed at biochemical problem solving, experimental design and the application of quantitative methods are integral to the course.			Registration also required for Recitation (see below)
BIOL 204-201	Biochemistry	Philip Arthur Rea Shannon Nicole Sturgeon	DRLB A5	T 03:00 PM-04:00 PM	BIOL 204 examines the basic principles of protein structure, protein purification and characterization, proteomics, enzyme kinetics and mechanism, membrane structure and function, metabolism, and cellular energy transduction. The primary objective is to provide life scientists with an appreciation of basic principles of modern biochemistry, and of how the current conceptual and technical framework arose. Emphasis is placed on the experimental approaches and reasoning behind the dissection and reconstitution of these processes in a biological and, in some cases, clinical context. Discussions directed at biochemical problem solving, experimental design and the application of quantitative methods are integral to the course.			Registration also required for Lecture (see below)
BIOL 204-202	Biochemistry	Philip Arthur Rea Shannon Nicole Sturgeon	DRLB A5	T 04:00 PM-05:00 PM	BIOL 204 examines the basic principles of protein structure, protein purification and characterization, proteomics, enzyme kinetics and mechanism, membrane structure and function, metabolism, and cellular energy transduction. The primary objective is to provide life scientists with an appreciation of basic principles of modern biochemistry, and of how the current conceptual and technical framework arose. Emphasis is placed on the experimental approaches and reasoning behind the dissection and reconstitution of these processes in a biological and, in some cases, clinical context. Discussions directed at biochemical problem solving, experimental design and the application of quantitative methods are integral to the course.			Registration also required for Lecture (see below)
BIOL 211-601	Essentials of Molec. Biol. & Genetics	John E. Zimmerman	GLAB 101	M 05:30 PM-08:20 PM	This course will survey the discipline of molecular genetics. Mendelian and molecular genetics will be discussed as well as the use of genetic analysis to address questions in all areas of biology. The processes of DNA replication, transcription, and translation will be discussed at the molecular level. Other topics include the regulation of gene expression and genomics. This course is open to students in the College of Liberal and Professional Studies only.			Course Must Be Taken For A Grade Registration also required for Recitation (see below)		https://pennintouchdaemon.apps.upenn.edu/pennInTouchProdDaemon/jsp/fast.do?webService=syllabus&term=2019A&course=BIOL211601
BIOL 211-602	Essentials of Molecular Biology and Genetics	John E. Zimmerman	GLAB 101	M 08:30 PM-09:00 PM	This course will survey the discipline of molecular genetics. Mendelian and molecular genetics will be discussed as well as the use of genetic analysis to address questions in all areas of biology. The processes of DNA replication, transcription, and translation will be discussed at the molecular level. Other topics include the regulation of gene expression and genomics. This course is open to students in the College of Liberal and Professional Studies only.			Registration also required for Lecture (see below)
BIOL 213-601	Essentials of Physiology	Xiaohong Zou Witmer	GLAB 100	TR 04:30 PM-05:50 PM	A comparative and quantitative approach to the physiological function of vertebrates. Topics include muscles, nervous system, cardiovascular system, respiration, and renal function. This course is open to students in the College of Liberal and Professional Studies only.			Course Must Be Taken For A Grade Registration also required for Recitation (see below)
BIOL 213-602	Essentials of Vertebrate Physiology		GLAB 100	TR 05:50 PM-06:20 PM	A comparative and quantitative approach to the physiological function of vertebrates. Topics include muscles, nervous system, cardiovascular system, respiration, and renal function. This course is open to students in the College of Liberal and Professional Studies only.			Registration also required for Lecture (see below)
BIOL 215-001	Vertebrate Physiology	Elizabeth Woodward	LLAB 10	TR 03:00 PM-04:30 PM	The course will focus on integrative aspects of physiological function of vertebrates. Comparative, environmental and quantitative approaches will be used. Major topics include muscle, the cardiovascular system, respiration, renal function and the nervous system.			Registration also required for Recitation (see below)
BIOL 215-201	Vertebrate Physiology	Cetewayo Saif Rashid	GLAB 101	T 04:30 PM-05:30 PM	The course will focus on integrative aspects of physiological function of vertebrates. Comparative, environmental and quantitative approaches will be used. Major topics include muscle, the cardiovascular system, respiration, renal function and the nervous system.			Registration also required for Lecture (see below)
BIOL 215-202	Vertebrate Physiology	Cetewayo Saif Rashid	GLAB 101	W 11:00 AM-12:00 PM	The course will focus on integrative aspects of physiological function of vertebrates. Comparative, environmental and quantitative approaches will be used. Major topics include muscle, the cardiovascular system, respiration, renal function and the nervous system.			Registration also required for Lecture (see below)
BIOL 215-203	Vertebrate Physiology	Arielle Richey Levine	GLAB 101	R 02:00 PM-03:00 PM	The course will focus on integrative aspects of physiological function of vertebrates. Comparative, environmental and quantitative approaches will be used. Major topics include muscle, the cardiovascular system, respiration, renal function and the nervous system.			Registration also required for Lecture (see below)
BIOL 215-204	Vertebrate Physiology	Arielle Richey Levine	GLAB 101	F 01:00 PM-02:00 PM	The course will focus on integrative aspects of physiological function of vertebrates. Comparative, environmental and quantitative approaches will be used. Major topics include muscle, the cardiovascular system, respiration, renal function and the nervous system.			Registration also required for Lecture (see below)
BIOL 221-401	Molec. Biol. & Genetics	Ishmail Abdus-Saboor Mia Tauna Levine	LEVN AUD	MWF 11:00 AM-12:00 PM	This course will survey the discipline of molecular genetics. Two broad areas will be considered 1) Molecular Biology: DNA replication, transcription, translation, regulation of gene expression in both prokaryotic and eukaryotic systems, and genomics and 2) Genetics: basic Mendelian & molecular genetics.	BIOL527401		Registration also required for Recitation (see below)
BIOL 221-402	Molecular Biology and Genetics	Abigail Marie Divito Joseph Zinski Priya Sivaramakrishnan	LLAB 10	R 05:00 PM-07:00 PM	This course will survey the discipline of molecular genetics. Two broad areas will be considered 1) Molecular Biology: DNA replication, transcription, translation, regulation of gene expression in both prokaryotic and eukaryotic systems, and genomics and 2) Genetics: basic Mendelian & molecular genetics.	BIOL527402		Registration also required for Lecture (see below)
BIOL 230-001	Evolutionary Biology	Paul Stutz Schmidt	LEVN AUD	TR 09:00 AM-10:30 AM	Theories and mechanisms of evolution, with emphasis on the genetic basis of evolutionary change.			Registration also required for Recitation (see below)
BIOL 230-201	Evolutionary Biology	Justin Troy Walsh	GLAB 100	T 03:00 PM-04:00 PM	Theories and mechanisms of evolution, with emphasis on the genetic basis of evolutionary change.			Registration also required for Lecture (see below)
BIOL 230-202	Evolutionary Biology	Ozan Kiratli	GLAB 100	W 10:00 AM-11:00 AM	Theories and mechanisms of evolution, with emphasis on the genetic basis of evolutionary change.			Registration also required for Lecture (see below)
BIOL 230-203	Evolutionary Biology	Ozan Kiratli	WILL 321	W 05:00 PM-06:00 PM	Theories and mechanisms of evolution, with emphasis on the genetic basis of evolutionary change.			Registration also required for Lecture (see below)
BIOL 230-204	Evolutionary Biology	Justin Troy Walsh	GLAB 100	R 03:00 PM-04:00 PM	Theories and mechanisms of evolution, with emphasis on the genetic basis of evolutionary change.			Registration also required for Lecture (see below)
BIOL 230-205	Evolutionary Biology	Andrew Grady Johnson	GLAB 100	F 12:00 PM-01:00 PM	Theories and mechanisms of evolution, with emphasis on the genetic basis of evolutionary change.			Registration also required for Lecture (see below)
BIOL 230-206	Evolutionary Biology	Andrew Grady Johnson	LLAB 109	T 11:00 AM-12:00 PM	Theories and mechanisms of evolution, with emphasis on the genetic basis of evolutionary change.			Contact Dept Or Instructor For Classrm Info Registration also required for Lecture (see below)
BIOL 275-601	Elements of Microbiology	Kieran Dilks	GLAB 101	F 04:00 PM-05:30 PM	Microbiology plays a central role in diverse areas of human life such as infectious disease, ecology, and biotechnology. This course will cover aspects of modern microbiology with an emphasis on prokaryotic organisms. The topics will include basic aspects of microbial diversity, genetics, and pathogenesis as well as examples of applied microbiology. This course is open to students in the College of Liberal and Professional Studies only.			Course Must Be Taken For A Grade Registration also required for Laboratory (see below)		https://pennintouchdaemon.apps.upenn.edu/pennInTouchProdDaemon/jsp/fast.do?webService=syllabus&term=2019A&course=BIOL275601
BIOL 275-602	Elements of Microbiology	Kieran Dilks	LEVN L11	F 05:30 PM-08:30 PM	Microbiology plays a central role in diverse areas of human life such as infectious disease, ecology, and biotechnology. This course will cover aspects of modern microbiology with an emphasis on prokaryotic organisms. The topics will include basic aspects of microbial diversity, genetics, and pathogenesis as well as examples of applied microbiology. This course is open to students in the College of Liberal and Professional Studies only.			Registration also required for Lecture (see below)
BIOL 275-603	Elements of Microbiology	Jessica A Ardis	LEVN L12	F 05:30 PM-08:30 PM	Microbiology plays a central role in diverse areas of human life such as infectious disease, ecology, and biotechnology. This course will cover aspects of modern microbiology with an emphasis on prokaryotic organisms. The topics will include basic aspects of microbial diversity, genetics, and pathogenesis as well as examples of applied microbiology. This course is open to students in the College of Liberal and Professional Studies only.			Only Open To LPS Pb Pre-Health Students Registration also required for Lecture (see below)
BIOL 304-601	Infectious Disease Bio	Nataliya Balashova	GLAB 100	R 06:30 PM-09:30 PM	This course focuses on selected topics concerning infectious agents, the diseases they cause in humans, and the social and scientific challenges they pose. The first section addresses the principles of epidemiology and microbial pathogenesis, as well as pathophysiology of infectious diseases. In the second section, tools and techniques of diagnosis, tracking, and control of infectious diseases will be discussed. To develop a broad understanding of the many different aspects of infectious processes, selected viral, fungal, protozoan, and helminthic pathogens and related infectious diseases will be presented. This course is open to students in the College of Liberal and Professional Studies only.			Only Open To LPS Pb Pre-Health Students		https://pennintouchdaemon.apps.upenn.edu/pennInTouchProdDaemon/jsp/fast.do?webService=syllabus&term=2019A&course=BIOL304601
BIOL 306-601	Histology	Deborah E Weil	LEVN L12	R 04:30 PM-07:30 PM	This course is designed to introduce the undergraduate student to the structure of tissues at the cellular level and to the way in which those tissues are assembled into organs. This knowledge of structure will be the basis for discussion of tissue and organ function. This course is open to students in the College of Liberal and Professional Studies only.			Only Open To LPS Pb Pre-Health Students Course Must Be Taken For A Grade
BIOL 330-001	Comp Vert Anatomy & Evol	Arthur E. Dunham	LLAB 109	MWF 10:00 AM-11:00 AM	This course will survey the phylogeny and anatomy of vertebrate organisms from a comparative evolutionary perspective. The lecture will concentrate on the history, diversity, structure and function of vertebrates. A companion lab course, BIOL 336, is available for those students interested in a more complete understanding of vertebrate anatomy.
BIOL 336-101	Comp Vert Anatomy Lab	Damien Lekkas	LLAB 117	T 06:00 PM-09:00 PM	Laboratory portion of BIOL 330 Comparative Vertebrate Anatomy and Evolution. Students will learn comparative anatomy through dissection of representative vertebrates. Students taking the lab must have credit or register for the lecture course, BIOL 330.
BIOL 354-601	Developmental Biology	John D. O. Wagner	LLAB 109	MW 05:00 PM-06:20 PM	A view of how an animal embryo is specified to develop and differentiate into a wide spectrum of cell types, and how the spatial patterns and axes of embyros are determined. The course will focus on genetic and molecular approaches, but will also cover the comparative anatomy of developing embryos to the extent necessary to understand the conserved aspects of embryonic patterning. Special emphasis will be placed on organisms with particular advantages for the study of embryonic development: e.g., mouse, frog, zebrafish, and Drosophila. The first half of the course will cover cell fate restrictions, cloning animals using nuclear transfer, stem cell biology, formation of the embryonic axes in vertebrates and Drosophila, and patterning of the neural tube and mesodermal tissues. The second half of the course will focus on emerging ideas and findings in the field, with emphasis on analysis of original literature.			Course Must Be Taken For A Grade Registration also required for Recitation (see below)
BIOL 354-602	Developmental Biology	John D. O. Wagner	LLAB 109	MW 06:30 PM-07:00 PM	A view of how an animal embryo is specified to develop and differentiate into a wide spectrum of cell types, and how the spatial patterns and axes of embyros are determined. The course will focus on genetic and molecular approaches, but will also cover the comparative anatomy of developing embryos to the extent necessary to understand the conserved aspects of embryonic patterning. Special emphasis will be placed on organisms with particular advantages for the study of embryonic development: e.g., mouse, frog, zebrafish, and Drosophila. The first half of the course will cover cell fate restrictions, cloning animals using nuclear transfer, stem cell biology, formation of the embryonic axes in vertebrates and Drosophila, and patterning of the neural tube and mesodermal tissues. The second half of the course will focus on emerging ideas and findings in the field, with emphasis on analysis of original literature.			Registration also required for Lecture (see below)
BIOL 375-401	Microbe Diversty&Pathogn: Microbial Diversity & Pathogenesis	Mechthild Pohlschroder	LEVN 111	TR 12:00 PM-01:30 PM	Microbiology plays a central role in diverse areas of human life such as infectious disease, ecology, and biotechnology. This course will cover aspects of modern microbiology with an emphasis on prokaryotic organisms. The topics will include basic aspects of microbial diversity, genetics, virology, and pathogenesis as well as examples of applied microbiology.	BIOL575401
BIOL 376-401	Microbial Diversity & Pathogenesis Lab	Karl G Siegert Byron P Sherwood	LEVN L57	TR 09:00 AM-12:00 PM	The importance of microbiology in complex issues, such as the impact of the microbiome in human health or as alternative energy sources, is being appreciated more and more each day. This upper level laboratory course provides students with a robust technical skill set while also giving them an opportunity to participate in an authentic research project that may lead to novel discoveries. Students will generate research questions, formulate hypotheses, design experiments, analyze data, and present their research findings to the class. In each project, students will use the cutting edge approach of metagenomics to evaluate the microbial diversity of their environment via Next Generation Sequencing. Students will also examine the function of microbial species within their communities. Potential projects include the isolation of novel antibiotic producers and the antibiotic they produce, designing and optimizing microbial fuel cells that can be used to generate electricity, or isolating antibiotic resistant bacteria and attempting novel approaches to inhibit or prevent their growth.	BIOL576401
BIOL 404-601	Immunobiology	Michael P Cancro	LEVN AUD	M 06:30 PM-09:30 PM	Early development of microbiology, pathology, and immunobiology; molecular and cellular bases of immune phenomena including: immunity to pathogens, immune diseases, autoimmunity, and hypersensitivity. This course is open to students in the College of Liberal and Professional Studies only.			Course Must Be Taken For A Grade
BIOL 406-301	Molecular Mechanisms of Infectious Disease Biology	David S. Roos		W 03:30 PM-06:30 PM	This course is designed for advanced undergraduates and beginning graduate students with a particular interest in infectious disease biology. Note that this course is not a comprehensive survey of the field and is not appropriate for students seeking a lecture course on disease. The primary objective of this course is to teach students considering a career in the biomedical sciences how to read, discuss, and question research papers effectively. Intensive classroom discussions focus on the experimental methods used, results obtained, interpretation of these results in the context of pathogen interactions with host cells and organisms, and implications for basic research and therapeutic development.			Contact Dept Or Instructor For Classrm Info
BIOL 407-601	Cancer Cell Biology	Shujuan Xia	GLAB 101	W 06:30 PM-09:30 PM	This course will focus on the molecular mechanisms by which fundamental cellular processes are disrupted in the development of cancer.			Course Must Be Taken For A Grade		https://pennintouchdaemon.apps.upenn.edu/pennInTouchProdDaemon/jsp/fast.do?webService=syllabus&term=2019A&course=BIOL407601
BIOL 415-401	Freshwater Ecology	Melinda Daniels David B. Arscott	LLAB 109	MW 03:30 PM-05:00 PM	Survey of the physical, chemical and biological properties of freshwater ecosystems, both riverine and lentic, natural and polluted.	ENVS416401
BIOL 417-301	Theoretical Population Biology	Erol Akcay	GLAB 100	MW 02:00 PM-03:30 PM	Introduction to basic theoretical tools to study the evolutionary and ecological dynamics of populations. Topics to be discussed include: basic population dynamics and population genetics theory, evolutionary game theory/adaptive dynamics, social evolution (kin selection/multilevel selection), life-history evolution, and stochastic models. Other topics may be added based on the specific interests of students in the class.
BIOL 425-101	Biochemistry and Molecular Genetics Superlab	John D. O. Wagner	LEVN L57	TR 01:30 PM-04:30 PM	Intensive laboratory class where open-ended, interesting biological problems are explored using modern lab techniques. Topics may include protein structure/function studies; genetic screens, genomics and gene expression studies; proteomics and protein purification techniques; and molecular cloning and DNA manipulation. The course emphasizes developing scientific communication and independent research skills. Course topics reflect the interests of individual Biology faculty members. This course is recommended for students considering independent research.			Communication Within the Curriculum
BIOL 431-401	Genome Sciences and Genomic Medicine	Brian D. Gregory	LEVN 111	TR 01:30 PM-03:00 PM	This course will be a focused study of genomes, genomic techniques, and how these approaches are and will be used in diagnosing and treating human disease. Topics will include genome sequencing, analysis of sequences and microarrays, and new techniques including high-throughput sequencing and reverse genetic analysis with a focus on genome-wide mutant collections.	CAMB431401
BIOL 433-301	Genetics of Adaptation:How Sex, Conflict & Pathogens Shape Modern Genomes	Mia Tauna Levine	COHN 337	T 03:00 PM-06:00 PM	In this course we explore the genetic basis of adaptation. We will investigate the forces that drive adaptation (e.g., environmental stress, pathogens, conflict, sex), the genetic mechanisms of adaptation (protein sequence changes, expression divergence, gene duplication, etc.), and the consequences of adaptation for contemporary cellular functions, developmental processes, and ecological interactions. The class meetings will be structured around both lectures and student-led discussions of the primary literature.
BIOL 438-301	Systems Biol: Int Phys	Lawrence C. Rome	MCNB 103	TR 01:30 PM-03:00 PM	The course will focus on muscle function from the level of molecules to whole animal locomotion. At each level of organization, muscle function will be explored from mechanical and energetic viewpoints. The course will include lectures, demonstrations, and several guest expert lectures. Students will also be introduced to realistic musculo-skeletal modelling and forward dynamic simulations to explore integrated function.
BIOL 456-301	Neural Circuits For Survival	J. Nicholas Betley	GLAB 100	W 03:30 PM-06:30 PM	A fundamental goal of neuroscience is to understand how neural circuits in the brain function to influence behavior. The aim of this course is to highlight the neural basis of behavior and discuss modern approaches and novel methods to study the neuronal control of classically studied aspects of behavior. Through a combination of discussions, student presentations, and interactive lectures, we will explore the neural systems that regulate the interactions an animal has with the external world. We will explore sensory systems (such as vision, taste, and olfaction), motor systems, and survival behaviors (such as feeding, drinking, mating, and aggression). The course evaluation will be based largely on written work, participation, and presentations.
BIOL 464-101	Field Studies in Aquatic Microbial Ecology	Byron P Sherwood	LEVN L57	F 11:00 AM-04:00 PM	This is a field and laboratory-based course that involves students in hands-on research methods in aquatic microbial ecology. Students will gain familiarity with experimental design, field measurements of environmental parameters (physical, chemical, and biological), sample collection techniques, and laboratory analyses required to assess the activity, health, and community composition of aquatic microbial ecosystems in an urban environment. We will compare and contrast various watersheds in and around the greater Philadelphia area, and students will design and conduct original independent research as a final course project.
BIOL 469-301	Plant Physiology Through Space and Time	Brent R. Helliker	LEVN L12	F 12:00 PM-03:00 PM	This course is a lab/lecture/seminar hybrid that will meet once per week for three hours. Each session will consist of mini-lecture/lab, paper discussions/lab, or solely lab efforts. All reading assignments will be available on Canvas (no textbook fees). We will exam various aspects of photosynthesis, water relations and nutrient acquisition in the context of the evolutionary progression of higher plants. With each subject, we will consider, measure, and in some cases model whole-plant physiology while examining sub-cellular-level controls and ecosystem-to-global-level consequences. This course is designed to give molecular biologists through earth-system scientists the tools to measure and understand whole-plant physiological responses to molecular manipulation and environmental variability. All students will learn to appreciate the context of their work on both micro and macro scales.
BIOL 480-401	Advanced Cell Biology	Wei Guo	GLAB 100	TR 10:30 AM-12:00 PM	This course is designed for beginning graduate students and advanced undergraduates with a particular enthusiasm for cell biology. Biology 480 does not attempt to cover all aspects of cell biology, and is therefore not appropriate for students seeking a lecture course which provides a comprehensive survey of the field. Rather, the primary objective of this course is to teach those students considering a career in the biomedical sciences how to read, discuss, and question original research papers effectively. Intensive classroom discussions focus on the experimental methods used, results obtained, interpretation of these results in the context of cell structure and function, and implications for further studies.	CAMB480401
BIOL 480-601	Advanced Cell Biology	Xiaohong Zou Witmer	GLAB 101	TR 07:00 PM-08:30 PM	This course is designed for beginning graduate students and advanced undergraduates with a particular enthusiasm for cell biology. Biology 480 does not attempt to cover all aspects of cell biology, and is therefore not appropriate for students seeking a lecture course which provides a comprehensive survey of the field. Rather, the primary objective of this course is to teach those students considering a career in the biomedical sciences how to read, discuss, and question original research papers effectively. Intensive classroom discussions focus on the experimental methods used, results obtained, interpretation of these results in the context of cell structure and function, and implications for further studies.			Only Open To LPS Pb Pre-Health Students Course Must Be Taken For A Grade
BIOL 527-401	Genetics For Comp Bio	Ishmail Abdus-Saboor Mia Tauna Levine	LEVN AUD	MWF 11:00 AM-12:00 PM	This course will survey the discipline of molecular genetics. Two broad areas will be considered: 1) Molecular biology: DNA replication, transcription, translation, and the regulation of gene expression in both prokaryotic and eukaryotic systems and genomics and 2) Genetics: basic Mendelian & molecular genetics.	BIOL221401		Permission Needed From Department Registration also required for Recitation (see below)
BIOL 527-402	Genetics For Computational Biology	Abigail Marie Divito Joseph Zinski Priya Sivaramakrishnan	LLAB 10	R 05:00 PM-07:00 PM	This course will survey the discipline of molecular genetics. Two broad areas will be considered: 1) Molecular biology: DNA replication, transcription, translation, and the regulation of gene expression in both prokaryotic and eukaryotic systems and genomics and 2) Genetics: basic Mendelian & molecular genetics.	BIOL221402		Permission Needed From Department Registration also required for Lecture (see below)
BIOL 537-401	Adv. Computational Biol	Yoseph Barash Kai Tan Christopher D. Brown Li-San Wang	BRB 252	TR 01:30 PM-03:00 PM	Discussion of special research topics.	GCB537401		Permission Needed From Department
BIOL 575-401	Microbe Diversty&Pathogn: Microbial Diversity & Pathogenesis	Mechthild Pohlschroder	LEVN 111	TR 12:00 PM-01:30 PM	Advanced version of BIOL 375: Microbial Diversity and Pathogenesis for graduate students only. Additional readings and course work as directed.	BIOL375401		Permission Needed From Department
BIOL 576-401	Microbial Diversity & Pathogenesis Lab	Karl G Siegert Byron P Sherwood	LEVN L57	TR 09:00 AM-12:00 PM	Advanced version of BIOL 376: Microbial Diversity and Pathogenesis Lab for graduate students only. Additional readings and course work as directed.	BIOL376401		Permission Needed From Department