Title	Instructors	Location	Time	Description	Cross listings	Fulfills	Registration notes	Syllabus	Syllabus URL
BIOL 1101-001	Introduction to Biology A	Jennifer E Round Lori Haynes Spindler John D Wagner	LLAB 10	MW 12:00 PM-1:29 PM	General principles of biology focusing on the basic chemistry of life, cell biology, molecular biology, and genetics in all types of living organisms. Particular emphasis will be given to links between the fundamental processes covered and current challenges of humankind in the areas of energy, food, and health.		Living World Sector
BIOL 1101-101	Introduction to Biology A	Linda Robinson Lori Haynes Spindler	LLAB 114	M 1:45 PM-4:44 PM	General principles of biology focusing on the basic chemistry of life, cell biology, molecular biology, and genetics in all types of living organisms. Particular emphasis will be given to links between the fundamental processes covered and current challenges of humankind in the areas of energy, food, and health.		Living World Sector
BIOL 1101-102	Introduction to Biology A	Linda Robinson Eunnuri Yi	LLAB 115	M 1:45 PM-4:44 PM	General principles of biology focusing on the basic chemistry of life, cell biology, molecular biology, and genetics in all types of living organisms. Particular emphasis will be given to links between the fundamental processes covered and current challenges of humankind in the areas of energy, food, and health.		Living World Sector
BIOL 1101-103	Introduction to Biology A	Linda Robinson Terrence Nadir Topping-Brown	LLAB 114	T 10:15 AM-1:14 PM	General principles of biology focusing on the basic chemistry of life, cell biology, molecular biology, and genetics in all types of living organisms. Particular emphasis will be given to links between the fundamental processes covered and current challenges of humankind in the areas of energy, food, and health.		Living World Sector
BIOL 1101-104	Introduction to Biology A	Linda Robinson Eunnuri Yi	LLAB 115	T 10:15 AM-1:14 PM	General principles of biology focusing on the basic chemistry of life, cell biology, molecular biology, and genetics in all types of living organisms. Particular emphasis will be given to links between the fundamental processes covered and current challenges of humankind in the areas of energy, food, and health.		Living World Sector
BIOL 1101-105	Introduction to Biology A	Linda Robinson Lori Haynes Spindler	LLAB 114	T 1:45 PM-4:44 PM	General principles of biology focusing on the basic chemistry of life, cell biology, molecular biology, and genetics in all types of living organisms. Particular emphasis will be given to links between the fundamental processes covered and current challenges of humankind in the areas of energy, food, and health.		Living World Sector
BIOL 1101-106	Introduction to Biology A	Isabella Joy Higgins Linda Robinson	LLAB 115	T 1:45 PM-4:44 PM	General principles of biology focusing on the basic chemistry of life, cell biology, molecular biology, and genetics in all types of living organisms. Particular emphasis will be given to links between the fundamental processes covered and current challenges of humankind in the areas of energy, food, and health.		Living World Sector
BIOL 1101-107	Introduction to Biology A	Linda Robinson Caitlin Tilsed	LLAB 114	W 8:30 AM-11:29 AM	General principles of biology focusing on the basic chemistry of life, cell biology, molecular biology, and genetics in all types of living organisms. Particular emphasis will be given to links between the fundamental processes covered and current challenges of humankind in the areas of energy, food, and health.		Living World Sector
BIOL 1101-108	Introduction to Biology A	Linda Robinson Lori Haynes Spindler	LLAB 114	W 1:45 PM-4:44 PM	General principles of biology focusing on the basic chemistry of life, cell biology, molecular biology, and genetics in all types of living organisms. Particular emphasis will be given to links between the fundamental processes covered and current challenges of humankind in the areas of energy, food, and health.		Living World Sector
BIOL 1101-109	Introduction to Biology A	Staver Bezhani Linda Robinson	LLAB 115	W 1:45 PM-4:44 PM	General principles of biology focusing on the basic chemistry of life, cell biology, molecular biology, and genetics in all types of living organisms. Particular emphasis will be given to links between the fundamental processes covered and current challenges of humankind in the areas of energy, food, and health.		Living World Sector
BIOL 1101-110	Introduction to Biology A	Linda Robinson Terrence Nadir Topping-Brown	LLAB 114	R 10:15 AM-1:14 PM	General principles of biology focusing on the basic chemistry of life, cell biology, molecular biology, and genetics in all types of living organisms. Particular emphasis will be given to links between the fundamental processes covered and current challenges of humankind in the areas of energy, food, and health.		Living World Sector
BIOL 1101-111	Introduction to Biology A	Linda Robinson Caitlin Tilsed	LLAB 115	R 10:15 AM-1:14 PM	General principles of biology focusing on the basic chemistry of life, cell biology, molecular biology, and genetics in all types of living organisms. Particular emphasis will be given to links between the fundamental processes covered and current challenges of humankind in the areas of energy, food, and health.		Living World Sector
BIOL 1101-112	Introduction to Biology A	Staver Bezhani Linda Robinson	LLAB 114	R 1:45 PM-4:44 PM	General principles of biology focusing on the basic chemistry of life, cell biology, molecular biology, and genetics in all types of living organisms. Particular emphasis will be given to links between the fundamental processes covered and current challenges of humankind in the areas of energy, food, and health.		Living World Sector
BIOL 1101-113	Introduction to Biology A	Isabella Joy Higgins Linda Robinson	LLAB 115	R 1:45 PM-4:44 PM	General principles of biology focusing on the basic chemistry of life, cell biology, molecular biology, and genetics in all types of living organisms. Particular emphasis will be given to links between the fundamental processes covered and current challenges of humankind in the areas of energy, food, and health.		Living World Sector
BIOL 1101-114	Introduction to Biology A	Linda Robinson Lori Haynes Spindler	LLAB 115	F 1:45 PM-4:44 PM	General principles of biology focusing on the basic chemistry of life, cell biology, molecular biology, and genetics in all types of living organisms. Particular emphasis will be given to links between the fundamental processes covered and current challenges of humankind in the areas of energy, food, and health.		Living World Sector
BIOL 1101-201	Introduction to Biology A	Jennifer E Round John D Wagner			General principles of biology focusing on the basic chemistry of life, cell biology, molecular biology, and genetics in all types of living organisms. Particular emphasis will be given to links between the fundamental processes covered and current challenges of humankind in the areas of energy, food, and health.		Living World Sector
BIOL 1101-601	Introduction to Biology A	Jessica A Ardis John E Zimmerman	LLAB 109	F 5:15 PM-8:14 PM	General principles of biology focusing on the basic chemistry of life, cell biology, molecular biology, and genetics in all types of living organisms. Particular emphasis will be given to links between the fundamental processes covered and current challenges of humankind in the areas of energy, food, and health.		Living World Sector
BIOL 1101-602	Introduction to Biology A	Jessica A Ardis Linda Robinson	LLAB 114	F 1:45 PM-4:44 PM	General principles of biology focusing on the basic chemistry of life, cell biology, molecular biology, and genetics in all types of living organisms. Particular emphasis will be given to links between the fundamental processes covered and current challenges of humankind in the areas of energy, food, and health.		Living World Sector
BIOL 1101-603	Introduction to Biology A	Jessica A Ardis Linda Robinson	LLAB 114	S 8:30 AM-11:29 AM	General principles of biology focusing on the basic chemistry of life, cell biology, molecular biology, and genetics in all types of living organisms. Particular emphasis will be given to links between the fundamental processes covered and current challenges of humankind in the areas of energy, food, and health.		Living World Sector
BIOL 1102-001	Introduction to Biology B	Dustin Brisson Katie Lynn Barott Sherwood	LEVN 111	MW 10:15 AM-11:44 AM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.		Living World Sector
BIOL 1102-101	Introduction to Biology B	Staver Bezhani Linda Robinson	LLAB 101	M 1:45 PM-4:44 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.		Living World Sector
BIOL 1102-102	Introduction to Biology B	Staver Bezhani Linda Robinson	LLAB 101	T 10:15 AM-1:14 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.		Living World Sector
BIOL 1102-103	Introduction to Biology B	Staver Bezhani Linda Robinson	LLAB 101	T 1:45 PM-4:44 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.		Living World Sector
BIOL 1102-104	Introduction to Biology B	Staver Bezhani Linda Robinson	LLAB 101	W 1:45 PM-4:44 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.		Living World Sector
BIOL 1110-401	Introduction to Brain and Behavior	Nicole C Rust	LEVN AUD	MW 12:00 PM-1:29 PM	Introduction to the structure and function of the vertebrate nervous system. We begin with the cellular basis of neuronal activities, then discuss the physiological bases of motor control, sensory systems, motivated behaviors, and higher mental processes. This course is intended for students interested in the neurobiology of behavior, ranging from animal behaviors to clinical disorders.	NRSC1110401, PSYC1210401	Living World Sector			https://coursesintouch.apps.upenn.edu/cpr/jsp/fast.do?webService=syll&t=202330&c=BIOL1110401
BIOL 1110-402	Introduction to Brain and Behavior	Mariela Lopez Valencia	LLAB 104	T 10:15 AM-11:44 AM	Introduction to the structure and function of the vertebrate nervous system. We begin with the cellular basis of neuronal activities, then discuss the physiological bases of motor control, sensory systems, motivated behaviors, and higher mental processes. This course is intended for students interested in the neurobiology of behavior, ranging from animal behaviors to clinical disorders.	NRSC1110402, PSYC1210402	Living World Sector
BIOL 1110-403	Introduction to Brain and Behavior	Joe Faryean	LLAB 104	T 12:00 PM-1:29 PM	Introduction to the structure and function of the vertebrate nervous system. We begin with the cellular basis of neuronal activities, then discuss the physiological bases of motor control, sensory systems, motivated behaviors, and higher mental processes. This course is intended for students interested in the neurobiology of behavior, ranging from animal behaviors to clinical disorders.	NRSC1110403, PSYC1210403	Living World Sector
BIOL 1110-404	Introduction to Brain and Behavior	Corinna Oswell	LLAB 104	T 1:45 PM-3:14 PM	Introduction to the structure and function of the vertebrate nervous system. We begin with the cellular basis of neuronal activities, then discuss the physiological bases of motor control, sensory systems, motivated behaviors, and higher mental processes. This course is intended for students interested in the neurobiology of behavior, ranging from animal behaviors to clinical disorders.	NRSC1110404, PSYC1210404	Living World Sector
BIOL 1110-405	Introduction to Brain and Behavior	Marc Carceles Cordon	LLAB 104	T 3:30 PM-4:59 PM	Introduction to the structure and function of the vertebrate nervous system. We begin with the cellular basis of neuronal activities, then discuss the physiological bases of motor control, sensory systems, motivated behaviors, and higher mental processes. This course is intended for students interested in the neurobiology of behavior, ranging from animal behaviors to clinical disorders.	NRSC1110405, PSYC1210405	Living World Sector
BIOL 1110-406	Introduction to Brain and Behavior	Angela Bongiovanni	LLAB 104	R 10:15 AM-11:44 AM	Introduction to the structure and function of the vertebrate nervous system. We begin with the cellular basis of neuronal activities, then discuss the physiological bases of motor control, sensory systems, motivated behaviors, and higher mental processes. This course is intended for students interested in the neurobiology of behavior, ranging from animal behaviors to clinical disorders.	NRSC1110406, PSYC1210406	Living World Sector
BIOL 1110-407	Introduction to Brain and Behavior	Ronni Kurzion	LLAB 104	R 12:00 PM-1:29 PM	Introduction to the structure and function of the vertebrate nervous system. We begin with the cellular basis of neuronal activities, then discuss the physiological bases of motor control, sensory systems, motivated behaviors, and higher mental processes. This course is intended for students interested in the neurobiology of behavior, ranging from animal behaviors to clinical disorders.	NRSC1110407, PSYC1210407	Living World Sector
BIOL 1110-408	Introduction to Brain and Behavior	Margaret Pecsok	LLAB 104	R 1:45 PM-3:14 PM	Introduction to the structure and function of the vertebrate nervous system. We begin with the cellular basis of neuronal activities, then discuss the physiological bases of motor control, sensory systems, motivated behaviors, and higher mental processes. This course is intended for students interested in the neurobiology of behavior, ranging from animal behaviors to clinical disorders.	NRSC1110408, PSYC1210408	Living World Sector
BIOL 1110-409	Introduction to Brain and Behavior	Aaron Mcknight	LLAB 104	R 3:30 PM-4:59 PM	Introduction to the structure and function of the vertebrate nervous system. We begin with the cellular basis of neuronal activities, then discuss the physiological bases of motor control, sensory systems, motivated behaviors, and higher mental processes. This course is intended for students interested in the neurobiology of behavior, ranging from animal behaviors to clinical disorders.	NRSC1110409, PSYC1210409	Living World Sector
BIOL 1110-601	Introduction to Brain and Behavior	Judith Mclean	GLAB 101	MW 5:15 PM-6:44 PM	Introduction to the structure and function of the vertebrate nervous system. We begin with the cellular basis of neuronal activities, then discuss the physiological bases of motor control, sensory systems, motivated behaviors, and higher mental processes. This course is intended for students interested in the neurobiology of behavior, ranging from animal behaviors to clinical disorders.	NRSC1110601, PSYC1210601	Living World Sector			https://coursesintouch.apps.upenn.edu/cpr/jsp/fast.do?webService=syll&t=202330&c=BIOL1110601
BIOL 1110-602	Introduction to Brain and Behavior	Judith Mclean	LLAB 104	T 5:15 PM-6:44 PM	Introduction to the structure and function of the vertebrate nervous system. We begin with the cellular basis of neuronal activities, then discuss the physiological bases of motor control, sensory systems, motivated behaviors, and higher mental processes. This course is intended for students interested in the neurobiology of behavior, ranging from animal behaviors to clinical disorders.	NRSC1110602, PSYC1210602	Living World Sector
BIOL 1110-603	Introduction to Brain and Behavior	Judith Mclean	LLAB 104	R 5:15 PM-6:44 PM	Introduction to the structure and function of the vertebrate nervous system. We begin with the cellular basis of neuronal activities, then discuss the physiological bases of motor control, sensory systems, motivated behaviors, and higher mental processes. This course is intended for students interested in the neurobiology of behavior, ranging from animal behaviors to clinical disorders.	NRSC1110603, PSYC1210603	Living World Sector
BIOL 1121-001	Introduction to Biology - The Molecular Biology of Life	Jessica A Ardis Mark D Goulian Michael A Lampson	LLAB 10	MW 10:15 AM-11:44 AM	An intensive introductory lecture course covering the cell, molecular biology, biochemistry, and the genetics of animals, bacteria, and viruses. This course is comparable to Biology 1101, but places greater emphasis on molecular mechanisms and experimental approaches. Particular attention is given to the ways in which modern cell biological and molecular genetic methods contribute to our understanding of evolutionary processes, the mechanistic basis of human disease, and recent biotechnological innovations. Students are encouraged to take BIOL 1121 and BIOL 1123 concurrently.		Quantitative Data Analysis Living World Sector			https://coursesintouch.apps.upenn.edu/cpr/jsp/fast.do?webService=syll&t=202330&c=BIOL1121001
BIOL 1121-201	Introduction to Biology - The Molecular Biology of Life	Jessica A Ardis Mark D Goulian Michael A Lampson	LLAB 10	F 10:15 AM-11:14 AM	An intensive introductory lecture course covering the cell, molecular biology, biochemistry, and the genetics of animals, bacteria, and viruses. This course is comparable to Biology 1101, but places greater emphasis on molecular mechanisms and experimental approaches. Particular attention is given to the ways in which modern cell biological and molecular genetic methods contribute to our understanding of evolutionary processes, the mechanistic basis of human disease, and recent biotechnological innovations. Students are encouraged to take BIOL 1121 and BIOL 1123 concurrently.		Quantitative Data Analysis Living World Sector			https://coursesintouch.apps.upenn.edu/cpr/jsp/fast.do?webService=syll&t=202330&c=BIOL1121201
BIOL 1123-001	Introductory Molecular Biology Laboratory	Jessica A Ardis Karl G Siegert	LLAB 10	F 8:30 AM-9:29 AM	An intensive introductory laboratory course emphasizing how molecular biology has revolutionized our understanding of cell and organism functions. BIOL 1121 and BIOL 1123 should be taken concurrently.
BIOL 1123-101	Introductory Molecular Biology Laboratory	Jessica A Ardis Karl G Siegert	LEVN L11	M 12:00 PM-2:59 PM	An intensive introductory laboratory course emphasizing how molecular biology has revolutionized our understanding of cell and organism functions. BIOL 1121 and BIOL 1123 should be taken concurrently.
BIOL 1123-102	Introductory Molecular Biology Laboratory	Jessica A Ardis Karl G Siegert		CANCELED	An intensive introductory laboratory course emphasizing how molecular biology has revolutionized our understanding of cell and organism functions. BIOL 1121 and BIOL 1123 should be taken concurrently.
BIOL 1123-103	Introductory Molecular Biology Laboratory	Jessica A Ardis Karl G Siegert	LEVN L11	T 10:15 AM-1:14 PM	An intensive introductory laboratory course emphasizing how molecular biology has revolutionized our understanding of cell and organism functions. BIOL 1121 and BIOL 1123 should be taken concurrently.
BIOL 1123-104	Introductory Molecular Biology Laboratory	Jessica A Ardis Karl G Siegert	LEVN L12	T 10:15 AM-1:14 PM	An intensive introductory laboratory course emphasizing how molecular biology has revolutionized our understanding of cell and organism functions. BIOL 1121 and BIOL 1123 should be taken concurrently.
BIOL 1123-105	Introductory Molecular Biology Laboratory	Jessica A Ardis Karl G Siegert	LEVN L11	T 3:30 PM-6:29 PM	An intensive introductory laboratory course emphasizing how molecular biology has revolutionized our understanding of cell and organism functions. BIOL 1121 and BIOL 1123 should be taken concurrently.
BIOL 1123-106	Introductory Molecular Biology Laboratory	Jessica A Ardis Karl G Siegert	LEVN L12	T 3:30 PM-6:29 PM	An intensive introductory laboratory course emphasizing how molecular biology has revolutionized our understanding of cell and organism functions. BIOL 1121 and BIOL 1123 should be taken concurrently.
BIOL 1123-107	Introductory Molecular Biology Laboratory	Jessica A Ardis Karl G Siegert	LEVN L11	W 1:45 PM-4:44 PM	An intensive introductory laboratory course emphasizing how molecular biology has revolutionized our understanding of cell and organism functions. BIOL 1121 and BIOL 1123 should be taken concurrently.
BIOL 1123-108	Introductory Molecular Biology Laboratory	Jessica A Ardis Karl G Siegert	LEVN L12	W 1:45 PM-4:44 PM	An intensive introductory laboratory course emphasizing how molecular biology has revolutionized our understanding of cell and organism functions. BIOL 1121 and BIOL 1123 should be taken concurrently.
BIOL 1123-109	Introductory Molecular Biology Laboratory	Jessica A Ardis Karl G Siegert	LEVN L11	R 8:30 AM-11:29 AM	An intensive introductory laboratory course emphasizing how molecular biology has revolutionized our understanding of cell and organism functions. BIOL 1121 and BIOL 1123 should be taken concurrently.
BIOL 1123-110	Introductory Molecular Biology Laboratory	Jessica A Ardis Karl G Siegert	LEVN L12	R 10:15 AM-1:14 PM	An intensive introductory laboratory course emphasizing how molecular biology has revolutionized our understanding of cell and organism functions. BIOL 1121 and BIOL 1123 should be taken concurrently.
BIOL 1123-111	Introductory Molecular Biology Laboratory	Jessica A Ardis Karl G Siegert	LEVN L11	R 12:00 PM-2:59 PM	An intensive introductory laboratory course emphasizing how molecular biology has revolutionized our understanding of cell and organism functions. BIOL 1121 and BIOL 1123 should be taken concurrently.
BIOL 1123-112	Introductory Molecular Biology Laboratory	Jessica A Ardis Karl G Siegert	LEVN L11	R 3:30 PM-6:29 PM	An intensive introductory laboratory course emphasizing how molecular biology has revolutionized our understanding of cell and organism functions. BIOL 1121 and BIOL 1123 should be taken concurrently.
BIOL 2001-601	Essentials of Cell Biology	Xiaohong Witmer	LLAB 109	TR 5:15 PM-6:44 PM	An intermediate level exploration of cell structure and function including membrane structure, intracellular organelles, membrane trafficking, surface receptors and signal transduction, the cytoskeleton, cell motility and communication, and the cell cycle. This course is open to students in the College of Liberal and Professional Studies only.
BIOL 2010-001	Cell Biology	Wei Guo Tatyana M Svitkina	LLAB 10	TR 10:15 AM-11:44 AM	A conceptual view of cell structure and function including membrane structure, intracellular organelles, membrane trafficking, surface receptors and signal transduction, the cytoskeleton, cell motility and communication, and the cell cycle. Cell biology is a dynamic field and recent research discoveries will be included in the lectures.
BIOL 2010-201	Cell Biology	Anna Palovcak	GLAB 101	T 5:15 PM-6:14 PM	A conceptual view of cell structure and function including membrane structure, intracellular organelles, membrane trafficking, surface receptors and signal transduction, the cytoskeleton, cell motility and communication, and the cell cycle. Cell biology is a dynamic field and recent research discoveries will be included in the lectures.
BIOL 2010-202	Cell Biology		GLAB 101	W 7:00 PM-7:59 PM	A conceptual view of cell structure and function including membrane structure, intracellular organelles, membrane trafficking, surface receptors and signal transduction, the cytoskeleton, cell motility and communication, and the cell cycle. Cell biology is a dynamic field and recent research discoveries will be included in the lectures.
BIOL 2110-401	Molecular and Cellular Neurobiology	Michael Kaplan Marc F Schmidt	LEVN AUD	MW 1:45 PM-3:14 PM	Cellular physiology of neurons and excitable cells; molecular neurobiology and development. Topics include: action potential generation; synaptic transmission; molecular and physiological studies of ion channels; second messengers; simple neural circuits; synaptic plasticity; learning and memory; and neural development.	NRSC2110401
BIOL 2110-402	Molecular and Cellular Neurobiology	Michael Kaplan	GLAB 102	W 7:00 PM-8:29 PM	Cellular physiology of neurons and excitable cells; molecular neurobiology and development. Topics include: action potential generation; synaptic transmission; molecular and physiological studies of ion channels; second messengers; simple neural circuits; synaptic plasticity; learning and memory; and neural development.	NRSC2110402
BIOL 2110-403	Molecular and Cellular Neurobiology	Michael Kaplan	GLAB 102	R 8:30 AM-9:59 AM	Cellular physiology of neurons and excitable cells; molecular neurobiology and development. Topics include: action potential generation; synaptic transmission; molecular and physiological studies of ion channels; second messengers; simple neural circuits; synaptic plasticity; learning and memory; and neural development.	NRSC2110403
BIOL 2110-404	Molecular and Cellular Neurobiology	Michael Kaplan	GLAB 102	R 1:45 PM-3:14 PM	Cellular physiology of neurons and excitable cells; molecular neurobiology and development. Topics include: action potential generation; synaptic transmission; molecular and physiological studies of ion channels; second messengers; simple neural circuits; synaptic plasticity; learning and memory; and neural development.	NRSC2110404
BIOL 2110-405	Molecular and Cellular Neurobiology	Michael Kaplan	GLAB 102	R 5:15 PM-6:44 PM	Cellular physiology of neurons and excitable cells; molecular neurobiology and development. Topics include: action potential generation; synaptic transmission; molecular and physiological studies of ion channels; second messengers; simple neural circuits; synaptic plasticity; learning and memory; and neural development.	NRSC2110405
BIOL 2110-406	Molecular and Cellular Neurobiology	Michael Kaplan	GLAB 102	F 10:15 AM-11:44 AM	Cellular physiology of neurons and excitable cells; molecular neurobiology and development. Topics include: action potential generation; synaptic transmission; molecular and physiological studies of ion channels; second messengers; simple neural circuits; synaptic plasticity; learning and memory; and neural development.	NRSC2110406
BIOL 2110-407	Molecular and Cellular Neurobiology	Michael Kaplan	GLAB 102	F 12:00 PM-1:29 PM	Cellular physiology of neurons and excitable cells; molecular neurobiology and development. Topics include: action potential generation; synaptic transmission; molecular and physiological studies of ion channels; second messengers; simple neural circuits; synaptic plasticity; learning and memory; and neural development.	NRSC2110407
BIOL 2110-408	Molecular and Cellular Neurobiology	Michael Kaplan	GLAB 102	F 3:30 PM-4:59 PM	Cellular physiology of neurons and excitable cells; molecular neurobiology and development. Topics include: action potential generation; synaptic transmission; molecular and physiological studies of ion channels; second messengers; simple neural circuits; synaptic plasticity; learning and memory; and neural development.	NRSC2110408
BIOL 2110-409	Molecular and Cellular Neurobiology	Michael Kaplan	GLAB 102	R 7:00 PM-8:29 PM	Cellular physiology of neurons and excitable cells; molecular neurobiology and development. Topics include: action potential generation; synaptic transmission; molecular and physiological studies of ion channels; second messengers; simple neural circuits; synaptic plasticity; learning and memory; and neural development.	NRSC2110409
BIOL 2140-401	Evolution of Behavior: Animal Behavior	Yun Ding Marc F Schmidt	LEVN AUD	TR 1:45 PM-3:14 PM	The evolution of behavior in animals will be explored using basic genetic and evolutionary principles. Lectures will highlight behavioral principles using a wide range of animal species, both vertebrate and invertebrate. Examples of behavior include the complex economic decisions related to foraging, migratory birds using geomagnetic fields to find breeding grounds, and the decision individuals make to live in groups. Group living has led to the evolution of social behavior and much of the course will focus on group formation, cooperation among kin, mating systems, territoriality and communication.	NRSC2140401, PSYC2220401
BIOL 2140-402	Evolution of Behavior: Animal Behavior		LEVN AUD	T 7:00 PM-7:59 PM	The evolution of behavior in animals will be explored using basic genetic and evolutionary principles. Lectures will highlight behavioral principles using a wide range of animal species, both vertebrate and invertebrate. Examples of behavior include the complex economic decisions related to foraging, migratory birds using geomagnetic fields to find breeding grounds, and the decision individuals make to live in groups. Group living has led to the evolution of social behavior and much of the course will focus on group formation, cooperation among kin, mating systems, territoriality and communication.	NRSC2140402, PSYC2220402
BIOL 2140-403	Evolution of Behavior: Animal Behavior		LLAB 109	F 10:15 AM-11:14 AM	The evolution of behavior in animals will be explored using basic genetic and evolutionary principles. Lectures will highlight behavioral principles using a wide range of animal species, both vertebrate and invertebrate. Examples of behavior include the complex economic decisions related to foraging, migratory birds using geomagnetic fields to find breeding grounds, and the decision individuals make to live in groups. Group living has led to the evolution of social behavior and much of the course will focus on group formation, cooperation among kin, mating systems, territoriality and communication.	NRSC2140403, PSYC2220403
BIOL 2201-601	Essentials of Molecular Biology and Genetics	John E Zimmerman	FAGN 218	M 5:15 PM-8:14 PM	This course will survey the discipline of molecular genetics. Mendelian and molecular genetics will be discussed as well as the use of genetic analysis to address questions in all areas of biology. The processes of DNA replication, transcription, and translation will be discussed at the molecular level. Other topics include the regulation of gene expression and genomics. This course is open to students in the College of Liberal and Professional Studies only.
BIOL 2210-401	Molecular Biology and Genetics	Nancy Bonini Kimberly L Gallagher	LLAB 10	TR 12:00 PM-1:29 PM	This course will survey the discipline of molecular genetics. Two broad areas will be considered 1) Molecular Biology: DNA replication, transcription, translation, regulation of gene expression in both prokaryotic and eukaryotic systems, and genomics and 2) Genetics: basic Mendelian & molecular genetics.	BIOL5210401				https://coursesintouch.apps.upenn.edu/cpr/jsp/fast.do?webService=syll&t=202330&c=BIOL2210401
BIOL 2210-402	Molecular Biology and Genetics		LEVN AUD	T 5:15 PM-6:14 PM	This course will survey the discipline of molecular genetics. Two broad areas will be considered 1) Molecular Biology: DNA replication, transcription, translation, regulation of gene expression in both prokaryotic and eukaryotic systems, and genomics and 2) Genetics: basic Mendelian & molecular genetics.	BIOL5210402
BIOL 2210-403	Molecular Biology and Genetics		FAGN 218	W 1:45 PM-2:44 PM	This course will survey the discipline of molecular genetics. Two broad areas will be considered 1) Molecular Biology: DNA replication, transcription, translation, regulation of gene expression in both prokaryotic and eukaryotic systems, and genomics and 2) Genetics: basic Mendelian & molecular genetics.	BIOL5210403
BIOL 2301-601	Essentials of Vertebrate Physiology	Xiaohong Witmer	GLAB 101	TR 7:00 PM-8:29 PM	A comparative and quantitative approach to the physiological function of vertebrates. Topics include muscles, nervous system, cardiovascular system, respiration, and renal function. This course is open to students in the College of Liberal and Professional Studies only.
BIOL 2510-401	Statistics for Biologists	Christopher Agard Sheel Chandra Aidan Fielding	LLAB 10	TR 8:30 AM-9:59 AM	Introductory probability theory. Principles of statistical methods. Problems of estimation and hypothesis testing in biology and related areas.	BIOL5510401
BIOL 2610-001	Ecology: From individuals to ecosystems	Erol Akcay Brent R Helliker	COHN 402	MW 12:00 PM-1:29 PM	The study of living organisms in their natural environment, spanning the ecological physiology of individuals, the structure of populations, and interactions among species, including the organization of communities and ecosystem function.
BIOL 2701-601	Elements of Microbiology	Jessica A Ardis Kieran Dilks	GLAB 101	F 5:15 PM-6:44 PM	Microbiology plays a central role in diverse areas of human life such as infectious disease, ecology, and biotechnology. This course will cover aspects of modern microbiology with an emphasis on prokaryotic organisms. The topics will include basic aspects of microbial diversity, genetics, and pathogenesis as well as examples of applied microbiology. This course is open to students in the College of Liberal and Professional Studies only.
BIOL 2701-602	Elements of Microbiology	Jessica A Ardis	LEVN L11	F 7:00 PM-9:59 PM	Microbiology plays a central role in diverse areas of human life such as infectious disease, ecology, and biotechnology. This course will cover aspects of modern microbiology with an emphasis on prokaryotic organisms. The topics will include basic aspects of microbial diversity, genetics, and pathogenesis as well as examples of applied microbiology. This course is open to students in the College of Liberal and Professional Studies only.
BIOL 2801-601	Essentials of Biochemistry	Ruth Elliott	FAGN 214	W 5:15 PM-8:14 PM	Intermediate level course covering principles of modern biochemistry. Topics include protein structure, protein purification and characterization, proteomics, enzyme kinetics and mechanisms, membrane structure and function, metabolism, and cellular energy transduction. Emphasis will be on biochemical problem solving, experimental design, and application of quantitative methods in a biological and clinical context. This course is open to students in the College of Liberal and Professional Studies only.
BIOL 3006-601	Histology	Brahim Chaqour	LEVN L12	R 5:15 PM-8:14 PM	This course is designed to introduce the undergraduate student to the structure of tissues at the cellular level and to the way in which those tissues are assembled into organs. This knowledge of structure will be the basis for discussion of tissue and organ function. This course is open to students in the College of Liberal and Professional Studies only.					https://coursesintouch.apps.upenn.edu/cpr/jsp/fast.do?webService=syll&t=202330&c=BIOL3006601
BIOL 3054-601	Developmental Biology	John D Wagner	LLAB 109	MW 5:15 PM-6:44 PM	A view of how an animal embryo is specified to develop and differentiate into a wide spectrum of cell types, and how the spatial patterns and axes of embyros are determined. The course will focus on genetic and molecular approaches, but will also cover the comparative anatomy of developing embryos to the extent necessary to understand the conserved aspects of embryonic patterning. Special emphasis will be placed on organisms with particular advantages for the study of embryonic development: e.g., mouse, frog, zebrafish, and Drosophila. The first half of the course will cover cell fate restrictions, cloning animals using nuclear transfer, stem cell biology, formation of the embryonic axes in vertebrates and Drosophila, and patterning of the neural tube and mesodermal tissues. The second half of the course will focus on emerging ideas and findings in the field, with emphasis on analysis of original literature.
BIOL 3310-001	Principles of Human Physiology	Yoichiro Mori	LEVN AUD	TR 12:00 PM-1:29 PM	Our focus will be on human physiology and we will cover most of the major organ systems in some depth. We seek to understand physiological phenomena using physical and chemical principles where possible. Basic cell and molecular biology, (bio)chemistry, physics and mathematics are prerequisites for the course, although we will quickly review the required background material when needed. Much of the motivation for the study of physiology is to understand disease, which in turn allows us to better appreciate normal physiology. We will discuss disease throughout the class. In physiology, structure often implies function, and we will thus also cover a fair amount of anatomy and histology.
BIOL 3310-201	Principles of Human Physiology	Yoichiro Mori	GLAB 100	T 5:15 PM-6:14 PM	Our focus will be on human physiology and we will cover most of the major organ systems in some depth. We seek to understand physiological phenomena using physical and chemical principles where possible. Basic cell and molecular biology, (bio)chemistry, physics and mathematics are prerequisites for the course, although we will quickly review the required background material when needed. Much of the motivation for the study of physiology is to understand disease, which in turn allows us to better appreciate normal physiology. We will discuss disease throughout the class. In physiology, structure often implies function, and we will thus also cover a fair amount of anatomy and histology.
BIOL 3310-202	Principles of Human Physiology	Yoichiro Mori	GLAB 100	W 8:30 AM-9:29 AM	Our focus will be on human physiology and we will cover most of the major organ systems in some depth. We seek to understand physiological phenomena using physical and chemical principles where possible. Basic cell and molecular biology, (bio)chemistry, physics and mathematics are prerequisites for the course, although we will quickly review the required background material when needed. Much of the motivation for the study of physiology is to understand disease, which in turn allows us to better appreciate normal physiology. We will discuss disease throughout the class. In physiology, structure often implies function, and we will thus also cover a fair amount of anatomy and histology.
BIOL 3310-203	Principles of Human Physiology	Yoichiro Mori	FAGN 110	W 3:30 PM-4:29 PM	Our focus will be on human physiology and we will cover most of the major organ systems in some depth. We seek to understand physiological phenomena using physical and chemical principles where possible. Basic cell and molecular biology, (bio)chemistry, physics and mathematics are prerequisites for the course, although we will quickly review the required background material when needed. Much of the motivation for the study of physiology is to understand disease, which in turn allows us to better appreciate normal physiology. We will discuss disease throughout the class. In physiology, structure often implies function, and we will thus also cover a fair amount of anatomy and histology.
BIOL 3310-204	Principles of Human Physiology	Yoichiro Mori	WILL 1	T 5:15 PM-6:14 PM	Our focus will be on human physiology and we will cover most of the major organ systems in some depth. We seek to understand physiological phenomena using physical and chemical principles where possible. Basic cell and molecular biology, (bio)chemistry, physics and mathematics are prerequisites for the course, although we will quickly review the required background material when needed. Much of the motivation for the study of physiology is to understand disease, which in turn allows us to better appreciate normal physiology. We will discuss disease throughout the class. In physiology, structure often implies function, and we will thus also cover a fair amount of anatomy and histology.
BIOL 4004-601	Immunobiology	Michael P Cancro Jennifer Londregan Jeffrey Maslanka	LEVN AUD	M 7:00 PM-9:59 PM	Early development of microbiology, pathology, and immunobiology; molecular and cellular bases of immune phenomena including: immunity to pathogens, immune diseases, autoimmunity, and hypersensitivity. This course is open to students in the College of Liberal and Professional Studies only.
BIOL 4007-601	Cancer Cell Biology	Maria Cotticelli Shujuan Xia	GLAB 100	W 7:00 PM-9:59 PM	This course will focus on the molecular mechanisms by which fundamental cellular processes are disrupted in the development of cancer.					https://coursesintouch.apps.upenn.edu/cpr/jsp/fast.do?webService=syll&t=202330&c=BIOL4007601
BIOL 4024-401	Cell Motility and the Cytoskeleton	Tatyana M Svitkina	GLAB 102	MW 3:30 PM-4:59 PM	Cytoskeleton and cell motility plays a crucial role in many aspects of normal and pathological physiology of individual cells, tissues, and whole organisms, including morphogenesis, immune response, wound healing, oncogenesis, and infection. This course will cover current topics in cell biology with emphasis on cytoskeleton and cell motility and their roles in these processes. Lectures, student presentations, and discussions in the class will be based on primary scientific literature.	BIOL5024401
BIOL 4142-401	Neurobiology of Learning and Memory	Mary Ellen Kelly	VAGL 2000	TR 1:45 PM-3:14 PM	This course focuses on the current state of our knowledge about the neurobiological basis of learning and memory. A combination of lectures and student seminars will explore the molecular and cellular basis of learning in invertebrates and vertebrates from a behavioral and neural perspective.	NRSC4442401, PSYC3301401				https://coursesintouch.apps.upenn.edu/cpr/jsp/fast.do?webService=syll&t=202330&c=BIOL4142401
BIOL 4266-401	Molecular Genetics of Neurological Disease	Nancy Bonini	LLAB 109	TR 10:15 AM-11:44 AM	This course will focus on the molecular basis of neurological diseases, exploring in detail key papers that cover topics including defining the disease genes, development of animal models that provide mechanistic insight, and seminal findings that reveal molecular understanding. Diseases covered will include neurological diseases of great focus today such as Alzheimer's, Fragile-X and autism, dementia, motor neuron degeneration, and microsatellite repeat expansion disorders. The course will provide a perspective from initial molecular determination through current status. Students will gain an understanding of how the molecular basis of a disease is discovered (from classical genetics to modern genomics) and how such diseases can be modeled in simple genetic systems for mechanistic insight. The course will be comprised of lectures with detailed analysis of primary literature and in-class activities. Grading will be based on class participation, exams, and written papers. Biology 2210 is a pre-requisite. Seniors are prioritized for the course	NRSC4266401				https://coursesintouch.apps.upenn.edu/cpr/jsp/fast.do?webService=syll&t=202330&c=BIOL4266401
BIOL 4310-301	Molecular Physiology	Dejian Ren	LLAB 109	MW 8:30 AM-9:59 AM	This course is designed for advanced undergraduate and graduate students who are interested in molecular physiology of sensory signal transduction. The major topics to cover will be signal transduction mechanisms used by membrane ion channels and receptors that detect the sensory stimuli (light, sound, temperature and taste, for example) and transmit the signals to the nervous system. Modern molecular/structural techniques (patch clamp, protein crystallization, molecular genetics, expression cloning and protein purification) will be introduced along with each topic. References will be primary research articles. Students will critically evaluate research discoveries by reading and presenting one to two original research papers. Each student is required to write a 10-page research proposal and to critique proposals written by fellow students.
BIOL 4430-401	Evolution and Ecology of Infectious Diseases	Dustin Brisson	GLAB 100	M 1:45 PM-4:44 PM	This course will focus on fundamental topics related to the ecological and evolutionary processes driving the transmission of pathogenic microbes among hosts including life-history strategies; evolution of pathogenic traits; the impacts of temporal, spatial and host-trait heterogeneity; and factors causing the emergence of an infectious pathogen. Examples will be drawn from human, wildlife, and plant pathogens to illustrate these ecological and evolutionary topics. Students will learn to develop and apply current ecological and evolutionary theory to infectious microbe research and gain practical experience accessing, interpreting and synthesizing the peer-reviewed scientific literature through a combination of popular and scientific readings, discussion, and lecture.	BIOL5430401
BIOL 4536-401	Introduction to Computational Biology & Biological Modeling	Gregory R Grant Junhyong Kim	LLAB 10	MW 3:30 PM-4:59 PM	The goal of this course is to develop a deeper understanding of techniques and concepts used in Computational Biology. The course will strive to focus on a small set of approaches to gain both theoretical and practical understanding of the methods. We will aim to cover practical issues such as programming and the use of programs, as well as theoretical issues such as algorithm design, statistical data analysis, theory of algorithms and statistics. This course WILL NOT provide a broad survey of the field nor teach specific tools but focus on a deep understanding of a small set of topics. We will discuss string algorithms, hidden markov models, dimension reduction, and machine learning (or phylogeny estimation) for biomedical problems.	BIOL5535401, CIS4360401	Natural Sciences & Mathematics Sector			https://coursesintouch.apps.upenn.edu/cpr/jsp/fast.do?webService=syll&t=202330&c=BIOL4536401
BIOL 4600-401	Field Botany	Timothy A Block Cynthia Skema			This course focuses on teaching students the Pennsylvania flora, both native and naturalized. Through weekly field trips, students will gain an appreciation for the diversity of plant species and plant communities in PA, and observe and discuss ecological and historical forces that influence plant species occurrences and plant communities. The ability to quickly and accurately identify plants in the field, through both sight identification and the use of a dichotomous key, is the major thrust of this course. Students will also learn how to appropriately collect plant materials for further study/identification in the laboratory and for archiving in an herbarium collection.	BIOL5600401
BIOL 4701-301	Prokaryotic Microbiology: A Pragmatic View	Mehmet Fevzi Daldal			This interactive course is intended for a small group of students aspiring to pursue research in microbiology, preferably using prokaryotes. Students will study selected papers and will attend the Prokaryotic Microbiology Seminars on Fridays. Specific problems of importance to a given field at a particular time will be critically analyzed and discussed: How were cutting edge techniques of the time used to address these problems? How would the same problems be approached using current techniques? The emphasis of the course will be on learning to become a thoughtful experimentalist rather than acquiring the hottest emerging knowledge.
BIOL 4825-401	Biochemistry and Molecular Genetics Superlab	Jennifer A Punt John D Wagner	LEVN L57	TR 1:45 PM-4:44 PM	Intensive laboratory class where open-ended, interesting biological problems are explored using modern lab techniques. Topics may include protein structure/function studies; genetic screens, genomics and gene expression studies; proteomics and protein purification techniques; and molecular cloning and DNA manipulation. The course emphasizes developing scientific communication and independent research skills. Course topics reflect the interests of individual Biology faculty members. This course is recommended for students considering independent research.	BIOL5825401
BIOL 5024-401	Cell Motility and the Cytoskeleton	Tatyana M Svitkina	GLAB 102	MW 3:30 PM-4:59 PM	Cytoskeleton and cell motility plays a crucial role in many aspects of normal and pathological physiology of individual cells, tissues, and whole organisms, including morphogenesis, immune response, wound healing, oncogenesis, and infection. This course will cover current topics in cell biology with emphasis on cytoskeleton and cell motility and their roles in these processes. Lectures, student presentations, and discussions in the class will be based on primary scientific literature.	BIOL4024401
BIOL 5210-401	Molecular Biology and Genetics	Nancy Bonini Kimberly L Gallagher	LLAB 10	TR 12:00 PM-1:29 PM	This course will survey the discipline of molecular genetics. Two broad areas will be considered 1) Molecular Biology: DNA replication, transcription, translation, regulation of gene expression in both prokaryotic and eukaryotic systems, and genomics and 2) Genetics: basic Mendelian & molecular genetics.	BIOL2210401
BIOL 5210-402	Molecular Biology and Genetics		LEVN AUD	T 5:15 PM-6:14 PM	This course will survey the discipline of molecular genetics. Two broad areas will be considered 1) Molecular Biology: DNA replication, transcription, translation, regulation of gene expression in both prokaryotic and eukaryotic systems, and genomics and 2) Genetics: basic Mendelian & molecular genetics.	BIOL2210402
BIOL 5210-403	Molecular Biology and Genetics		FAGN 218	W 1:45 PM-2:44 PM	This course will survey the discipline of molecular genetics. Two broad areas will be considered 1) Molecular Biology: DNA replication, transcription, translation, regulation of gene expression in both prokaryotic and eukaryotic systems, and genomics and 2) Genetics: basic Mendelian & molecular genetics.	BIOL2210403
BIOL 5430-401	Evolution and Ecology of Infectious Diseases	Dustin Brisson	GLAB 100	M 1:45 PM-4:44 PM	This course will focus on fundamental topics related to the ecological and evolutionary processes driving the transmission of pathogenic microbes among hosts including life-history strategies; evolution of pathogenic traits; the impacts of temporal, spatial and host-trait heterogeneity; and factors causing the emergence of an infectious pathogen. Examples will be drawn from human, wildlife, and plant pathogens to illustrate these ecological and evolutionary topics. Students will learn to develop and apply current ecological and evolutionary theory to infectious microbe research and gain practical experience accessing, interpreting and synthesizing the peer-reviewed scientific literature through a combination of popular and scientific readings, discussion, and lecture.	BIOL4430401
BIOL 5510-401	Statistics for Biologists	Christopher Agard Sheel Chandra Aidan Fielding	LLAB 10	TR 8:30 AM-9:59 AM	Introductory probability theory. Principles of statistical methods. Problems of estimation and hypothesis testing in biology and related areas.	BIOL2510401
BIOL 5535-401	Introduction to Computational Biology & Biological Modeling	Gregory R Grant Junhyong Kim	LLAB 10	MW 3:30 PM-4:59 PM	The goal of this course is to develop a deeper understanding of techniques and concepts used in Computational Biology. The course will strive to focus on a small set of approaches to gain both theoretical and practical understanding of the methods. We will aim to cover practical issues such as programming and the use of programs, as well as theoretical issues such as algorithm design, statistical data analysis, theory of algorithms and statistics. This course WILL NOT provide a broad survey of the field nor teach specific tools but focus on a deep understanding of a small set of topics. We will discuss string algorithms, hidden markov models, dimension reduction, and machine learning (or phylogeny estimation) for biomedical problems.	BIOL4536401, CIS4360401				https://coursesintouch.apps.upenn.edu/cpr/jsp/fast.do?webService=syll&t=202330&c=BIOL5535401
BIOL 5600-401	Field Botany	Timothy A Block Cynthia Skema			This course focuses on teaching students the Pennsylvania flora, both native and naturalized. Through weekly field trips, students will gain an appreciation for the diversity of plant species and plant communities in PA, and observe and discuss ecological and historical forces that influence plant species occurrences and plant communities. The ability to quickly and accurately identify plants in the field, through both sight identification and the use of a dichotomous key, is the major thrust of this course. Students will also learn how to appropriately collect plant materials for further study/identification in the laboratory and for archiving in an herbarium collection.	BIOL4600401
BIOL 5825-401	Biochemistry and Molecular Genetics Superlab	Jennifer A Punt John D Wagner	LEVN L57	TR 1:45 PM-4:44 PM	Intensive laboratory class where open-ended, interesting biological problems are explored using modern lab techniques. Topics may include protein structure/function studies; genetic screens, genomics and gene expression studies; proteomics and protein purification techniques; and molecular cloning and DNA manipulation. The course emphasizes developing scientific communication and independent research skills. Course topics reflect the interests of individual Biology faculty members. This course is recommended for students considering independent research.	BIOL4825401
BIOL 5860-401	Mathematical Modeling in Biology	Albane Thery	GLAB 101	MW 1:45 PM-3:14 PM	This course will cover various mathematical models and tools that are used to study modern biological problems. Mathematical models may be drawn from cell biology, physiology, population genetics, or ecology. Tools in dynamical systems or stochastic processes will be introduced as necessary. No prior knowledge of biology is needed to take this course, but some familiarity with differential equations and probability will be assumed.	MATH5861401				https://coursesintouch.apps.upenn.edu/cpr/jsp/fast.do?webService=syll&t=202330&c=BIOL5860401
BIOL 7000-301	Advanced Topics in Current Biological Research	David S Roos			Integrative seminar on current biological research for first-year PhD students.