Title	Instructors	Location	Time	Description	Cross listings	Fulfills	Registration notes	Syllabus	Syllabus URL
BIOL 1017-001	The Biology of Food	Richard Scott Poethig		MWF 12:00 PM-12:59 PM	This course will examine the ways in which humans manipulate - and have been manipulated by - the organisms we depend on for food, with particular emphasis on the biological factors that influence this interaction. The first part of the course will cover the biology, genetics, evolution, and breeding of cultivated plants and animals; the second part will concern the ecological, economic, and political factors that influence food production.		Natural Sciences & Mathematics Sector			https://coursesintouch.apps.upenn.edu/cpr/jsp/fast.do?webService=syll&t=202410&c=BIOL1017001
BIOL 1101-001	Introduction to Biology A	John Nicholas Betley Doris Wagner		MW 12:00 PM-1:29 PM	General principles of biology focusing on the basic chemistry of life, cell biology, molecular biology, and genetics in all types of living organisms. Particular emphasis will be given to links between the fundamental processes covered and current challenges of humankind in the areas of energy, food, and health.		Living World Sector
BIOL 1101-101	Introduction to Biology A	Staver Bezhani Linda Robinson		M 1:45 PM-4:44 PM	General principles of biology focusing on the basic chemistry of life, cell biology, molecular biology, and genetics in all types of living organisms. Particular emphasis will be given to links between the fundamental processes covered and current challenges of humankind in the areas of energy, food, and health.		Living World Sector
BIOL 1101-102	Introduction to Biology A	Staver Bezhani Linda Robinson		T 10:15 AM-1:14 PM	General principles of biology focusing on the basic chemistry of life, cell biology, molecular biology, and genetics in all types of living organisms. Particular emphasis will be given to links between the fundamental processes covered and current challenges of humankind in the areas of energy, food, and health.		Living World Sector
BIOL 1101-103	Introduction to Biology A	Staver Bezhani Linda Robinson		T 1:45 PM-4:44 PM	General principles of biology focusing on the basic chemistry of life, cell biology, molecular biology, and genetics in all types of living organisms. Particular emphasis will be given to links between the fundamental processes covered and current challenges of humankind in the areas of energy, food, and health.		Living World Sector
BIOL 1101-104	Introduction to Biology A	Staver Bezhani Linda Robinson		R 1:45 PM-4:44 PM	General principles of biology focusing on the basic chemistry of life, cell biology, molecular biology, and genetics in all types of living organisms. Particular emphasis will be given to links between the fundamental processes covered and current challenges of humankind in the areas of energy, food, and health.		Living World Sector
BIOL 1102-001	Introduction to Biology B	Lori Haynes Spindler		MW 12:00 PM-1:29 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.		Living World Sector
BIOL 1102-101	Introduction to Biology B	Linda Robinson		M 1:45 PM-4:44 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.		Living World Sector
BIOL 1102-102	Introduction to Biology B	Linda Robinson		M 1:45 PM-4:44 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.		Living World Sector
BIOL 1102-103	Introduction to Biology B	Linda Robinson		T 10:15 AM-1:14 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.		Living World Sector
BIOL 1102-104	Introduction to Biology B	Linda Robinson		T 10:15 AM-1:14 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.		Living World Sector
BIOL 1102-105	Introduction to Biology B	Linda Robinson		T 1:45 PM-4:44 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.		Living World Sector
BIOL 1102-106	Introduction to Biology B	Linda Robinson		T 1:45 PM-4:44 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.		Living World Sector
BIOL 1102-107	Introduction to Biology B	Linda Robinson		W 8:30 AM-11:29 AM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.		Living World Sector
BIOL 1102-108	Introduction to Biology B	Linda Robinson		W 1:45 PM-4:44 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.		Living World Sector
BIOL 1102-109	Introduction to Biology B	Linda Robinson		W 1:45 PM-4:44 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.		Living World Sector
BIOL 1102-110	Introduction to Biology B	Linda Robinson		R 10:15 AM-1:14 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.		Living World Sector
BIOL 1102-111	Introduction to Biology B	Linda Robinson		R 1:45 PM-4:44 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.		Living World Sector
BIOL 1102-112	Introduction to Biology B	Linda Robinson		R 1:45 PM-4:44 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.		Living World Sector
BIOL 1102-113	Introduction to Biology B	Linda Robinson		F 1:45 PM-4:44 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.		Living World Sector
BIOL 1102-601	Introduction to Biology B	John E. Zimmerman		F 5:15 PM-8:14 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.		Living World Sector
BIOL 1102-602	Introduction to Biology B	Linda Robinson		F 1:45 PM-4:44 PM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.		Living World Sector
BIOL 1102-603	Introduction to Biology B	Linda Robinson		S 8:30 AM-11:29 AM	General principles of biology focusing on evolution, physiology, development, and ecology in all types of living organisms.		Living World Sector
BIOL 1110-401	Introduction to Brain and Behavior	Michael Kane		TR 10:15 AM-11:44 AM	Introduction to the structure and function of the vertebrate nervous system. We begin with the cellular basis of neuronal activities, then discuss the physiological bases of motor control, sensory systems, motivated behaviors, and higher mental processes. This course is intended for students interested in the neurobiology of behavior, ranging from animal behaviors to clinical disorders.	NRSC1110401, PSYC1210401	Living World Sector
BIOL 1110-402	Introduction to Brain and Behavior	Venkata Sai Chaluvadi Michael Kane		M 8:30 AM-9:59 AM	Introduction to the structure and function of the vertebrate nervous system. We begin with the cellular basis of neuronal activities, then discuss the physiological bases of motor control, sensory systems, motivated behaviors, and higher mental processes. This course is intended for students interested in the neurobiology of behavior, ranging from animal behaviors to clinical disorders.	NRSC1110402, PSYC1210402	Living World Sector
BIOL 1110-403	Introduction to Brain and Behavior	Adriana Hernandez Vasquez Michael Kane		M 10:15 AM-11:44 AM	Introduction to the structure and function of the vertebrate nervous system. We begin with the cellular basis of neuronal activities, then discuss the physiological bases of motor control, sensory systems, motivated behaviors, and higher mental processes. This course is intended for students interested in the neurobiology of behavior, ranging from animal behaviors to clinical disorders.	NRSC1110403, PSYC1210403	Living World Sector
BIOL 1110-404	Introduction to Brain and Behavior	Michael Kane Marissa Maroni		M 12:00 PM-1:29 PM	Introduction to the structure and function of the vertebrate nervous system. We begin with the cellular basis of neuronal activities, then discuss the physiological bases of motor control, sensory systems, motivated behaviors, and higher mental processes. This course is intended for students interested in the neurobiology of behavior, ranging from animal behaviors to clinical disorders.	NRSC1110404, PSYC1210404	Living World Sector
BIOL 1110-405	Introduction to Brain and Behavior	Sierra Foshe Michael Kane		M 1:45 PM-3:14 PM	Introduction to the structure and function of the vertebrate nervous system. We begin with the cellular basis of neuronal activities, then discuss the physiological bases of motor control, sensory systems, motivated behaviors, and higher mental processes. This course is intended for students interested in the neurobiology of behavior, ranging from animal behaviors to clinical disorders.	NRSC1110405, PSYC1210405	Living World Sector
BIOL 1110-406	Introduction to Brain and Behavior	Sarah Applebey Michael Kane		M 3:30 PM-4:59 PM	Introduction to the structure and function of the vertebrate nervous system. We begin with the cellular basis of neuronal activities, then discuss the physiological bases of motor control, sensory systems, motivated behaviors, and higher mental processes. This course is intended for students interested in the neurobiology of behavior, ranging from animal behaviors to clinical disorders.	NRSC1110406, PSYC1210406	Living World Sector
BIOL 1110-407	Introduction to Brain and Behavior	Brianna Disanza Michael Kane		M 5:15 PM-6:44 PM	Introduction to the structure and function of the vertebrate nervous system. We begin with the cellular basis of neuronal activities, then discuss the physiological bases of motor control, sensory systems, motivated behaviors, and higher mental processes. This course is intended for students interested in the neurobiology of behavior, ranging from animal behaviors to clinical disorders.	NRSC1110407, PSYC1210407	Living World Sector
BIOL 1110-408	Introduction to Brain and Behavior	David Goldberg Michael Kane		M 7:00 PM-8:29 PM	Introduction to the structure and function of the vertebrate nervous system. We begin with the cellular basis of neuronal activities, then discuss the physiological bases of motor control, sensory systems, motivated behaviors, and higher mental processes. This course is intended for students interested in the neurobiology of behavior, ranging from animal behaviors to clinical disorders.	NRSC1110408, PSYC1210408	Living World Sector
BIOL 1110-409	Introduction to Brain and Behavior	Michael Kane Marcos Sanchez Navarro		T 8:30 AM-9:59 AM	Introduction to the structure and function of the vertebrate nervous system. We begin with the cellular basis of neuronal activities, then discuss the physiological bases of motor control, sensory systems, motivated behaviors, and higher mental processes. This course is intended for students interested in the neurobiology of behavior, ranging from animal behaviors to clinical disorders.	NRSC1110409, PSYC1210409	Living World Sector
BIOL 1110-601	Introduction to Brain and Behavior	Judith Mclean		TR 5:15 PM-6:44 PM	Introduction to the structure and function of the vertebrate nervous system. We begin with the cellular basis of neuronal activities, then discuss the physiological bases of motor control, sensory systems, motivated behaviors, and higher mental processes. This course is intended for students interested in the neurobiology of behavior, ranging from animal behaviors to clinical disorders.	NRSC1110601, PSYC1210601	Living World Sector
BIOL 1110-602	Introduction to Brain and Behavior	Judith Mclean		M 5:15 PM-6:44 PM	Introduction to the structure and function of the vertebrate nervous system. We begin with the cellular basis of neuronal activities, then discuss the physiological bases of motor control, sensory systems, motivated behaviors, and higher mental processes. This course is intended for students interested in the neurobiology of behavior, ranging from animal behaviors to clinical disorders.	NRSC1110602, PSYC1210602	Living World Sector
BIOL 1110-603	Introduction to Brain and Behavior	Judith Mclean		W 5:15 PM-6:44 PM	Introduction to the structure and function of the vertebrate nervous system. We begin with the cellular basis of neuronal activities, then discuss the physiological bases of motor control, sensory systems, motivated behaviors, and higher mental processes. This course is intended for students interested in the neurobiology of behavior, ranging from animal behaviors to clinical disorders.	NRSC1110603, PSYC1210603	Living World Sector
BIOL 1124-001	Introductory Organismal Biology Lab	Karl G Siegert		F 10:15 AM-11:14 AM	An intensive introductory laboratory course in organismal biology.
BIOL 1124-101	Introductory Organismal Biology Lab	Karl G Siegert		M 12:00 PM-2:59 PM	An intensive introductory laboratory course in organismal biology.
BIOL 1124-102	Introductory Organismal Biology Lab	Karl G Siegert		T 10:15 AM-1:14 PM	An intensive introductory laboratory course in organismal biology.
BIOL 1124-103	Introductory Organismal Biology Lab	Karl G Siegert		T 3:30 PM-6:29 PM	An intensive introductory laboratory course in organismal biology.
BIOL 1124-104	Introductory Organismal Biology Lab	Karl G Siegert		W 10:15 AM-1:14 PM	An intensive introductory laboratory course in organismal biology.
BIOL 1124-105	Introductory Organismal Biology Lab	Karl G Siegert		W 1:45 PM-4:44 PM	An intensive introductory laboratory course in organismal biology.
BIOL 1124-106	Introductory Organismal Biology Lab	Karl G Siegert		R 3:30 PM-6:29 PM	An intensive introductory laboratory course in organismal biology.
BIOL 1850-301	Research in Biological Sciences and its Social Impact (SNF Paideia Program Course)	Mechthild Pohlschroder			Working to remove the myths about fundamental and translational research, this course focuses on informing students beyond the public perception of biology and biological research. Striving to develop students' scientific communication skills, personal identity in science, and the intersection between research and community, we will engage students through collaboration with the Philadelphia community in addition to lecture and discussion based learning.
BIOL 2201-601	Essentials of Molecular Biology and Genetics	John E. Zimmerman			This course will survey the discipline of molecular genetics. Mendelian and molecular genetics will be discussed as well as the use of genetic analysis to address questions in all areas of biology. The processes of DNA replication, transcription, and translation will be discussed at the molecular level. Other topics include the regulation of gene expression and genomics. This course is open to students in the College of Liberal and Professional Studies only.
BIOL 2210-401	Molecular Biology and Genetics	Aman Yebio Husbands Mia Tauna Levine		TR 10:15 AM-11:44 AM	This course will survey the discipline of molecular genetics. Two broad areas will be considered 1) Molecular Biology: DNA replication, transcription, translation, regulation of gene expression in both prokaryotic and eukaryotic systems, and genomics and 2) Genetics: basic Mendelian & molecular genetics.	BIOL5210401
BIOL 2210-402	Molecular Biology and Genetics			W 12:00 PM-1:29 PM	This course will survey the discipline of molecular genetics. Two broad areas will be considered 1) Molecular Biology: DNA replication, transcription, translation, regulation of gene expression in both prokaryotic and eukaryotic systems, and genomics and 2) Genetics: basic Mendelian & molecular genetics.	BIOL5210402
BIOL 2210-403	Molecular Biology and Genetics			W 5:15 PM-6:44 PM	This course will survey the discipline of molecular genetics. Two broad areas will be considered 1) Molecular Biology: DNA replication, transcription, translation, regulation of gene expression in both prokaryotic and eukaryotic systems, and genomics and 2) Genetics: basic Mendelian & molecular genetics.	BIOL5210403
BIOL 2311-001	Human Physiology	Xiaohong Witmer		TR 8:30 AM-9:59 AM	This course examines the physiological mechanisms underlying homeostasis in humans. Integration from the cellular to organismal level as well as cooperation of multiple organ systems will be explored. Examples of pathophysiology during disease states will be discussed and highlighted. Although the focus will be on humans, we will study comparative aspects from other vertebrate and non-vertebrate organisms.
BIOL 2311-201	Human Physiology			T 12:00 PM-12:59 PM	This course examines the physiological mechanisms underlying homeostasis in humans. Integration from the cellular to organismal level as well as cooperation of multiple organ systems will be explored. Examples of pathophysiology during disease states will be discussed and highlighted. Although the focus will be on humans, we will study comparative aspects from other vertebrate and non-vertebrate organisms.
BIOL 2311-202	Human Physiology			F 10:15 AM-11:14 AM	This course examines the physiological mechanisms underlying homeostasis in humans. Integration from the cellular to organismal level as well as cooperation of multiple organ systems will be explored. Examples of pathophysiology during disease states will be discussed and highlighted. Although the focus will be on humans, we will study comparative aspects from other vertebrate and non-vertebrate organisms.
BIOL 2410-401	Evolutionary Biology	Paul Robert Schmidt Corlett Wood		TR 10:15 AM-11:44 AM	Theories and mechanisms of evolution, with emphasis on the genetic basis of evolutionary change.	BIOL5410401
BIOL 2410-402	Evolutionary Biology			T 12:00 PM-12:59 PM	Theories and mechanisms of evolution, with emphasis on the genetic basis of evolutionary change.
BIOL 2410-403	Evolutionary Biology			T 3:30 PM-4:29 PM	Theories and mechanisms of evolution, with emphasis on the genetic basis of evolutionary change.
BIOL 2410-404	Evolutionary Biology			W 10:15 AM-11:14 AM	Theories and mechanisms of evolution, with emphasis on the genetic basis of evolutionary change.
BIOL 2410-405	Evolutionary Biology			W 5:15 PM-6:14 PM	Theories and mechanisms of evolution, with emphasis on the genetic basis of evolutionary change.
BIOL 2410-406	Evolutionary Biology			R 12:00 PM-12:59 PM	Theories and mechanisms of evolution, with emphasis on the genetic basis of evolutionary change.	BIOL5410406
BIOL 2410-407	Evolutionary Biology			F 12:00 PM-12:59 PM	Theories and mechanisms of evolution, with emphasis on the genetic basis of evolutionary change.
BIOL 2701-601	Elements of Microbiology	Kieran Dilks		F 5:15 PM-6:44 PM	Microbiology plays a central role in diverse areas of human life such as infectious disease, ecology, and biotechnology. This course will cover aspects of modern microbiology with an emphasis on prokaryotic organisms. The topics will include basic aspects of microbial diversity, genetics, and pathogenesis as well as examples of applied microbiology. This course is open to students in the College of Liberal and Professional Studies only.
BIOL 2701-602	Elements of Microbiology	Jessica A Ardis		F 7:00 PM-9:59 PM	Microbiology plays a central role in diverse areas of human life such as infectious disease, ecology, and biotechnology. This course will cover aspects of modern microbiology with an emphasis on prokaryotic organisms. The topics will include basic aspects of microbial diversity, genetics, and pathogenesis as well as examples of applied microbiology. This course is open to students in the College of Liberal and Professional Studies only.
BIOL 2801-601	Essentials of Biochemistry	Ruth Elliott		W 5:15 PM-8:14 PM	Intermediate level course covering principles of modern biochemistry. Topics include protein structure, protein purification and characterization, proteomics, enzyme kinetics and mechanisms, membrane structure and function, metabolism, and cellular energy transduction. Emphasis will be on biochemical problem solving, experimental design, and application of quantitative methods in a biological and clinical context. This course is open to students in the College of Liberal and Professional Studies only.
BIOL 2810-001	Biochemistry	Philip Arthur Rea		MW 1:45 PM-3:14 PM	BIOL 204 examines the basic principles of protein structure, protein purification and characterization, proteomics, enzyme kinetics and mechanism, membrane structure and function, metabolism, and cellular energy transduction. The primary objective is to provide life scientists with an appreciation of basic principles of modern biochemistry, and of how the current conceptual and technical framework arose. Emphasis is placed on the experimental approaches and reasoning behind the dissection and reconstitution of these processes in a biological and, in some cases, clinical context. Discussions directed at biochemical problem solving, experimental design and the application of quantitative methods are integral to the course.
BIOL 2810-201	Biochemistry			T 7:00 PM-7:59 PM	BIOL 204 examines the basic principles of protein structure, protein purification and characterization, proteomics, enzyme kinetics and mechanism, membrane structure and function, metabolism, and cellular energy transduction. The primary objective is to provide life scientists with an appreciation of basic principles of modern biochemistry, and of how the current conceptual and technical framework arose. Emphasis is placed on the experimental approaches and reasoning behind the dissection and reconstitution of these processes in a biological and, in some cases, clinical context. Discussions directed at biochemical problem solving, experimental design and the application of quantitative methods are integral to the course.
BIOL 2810-202	Biochemistry			R 5:15 PM-6:14 PM	BIOL 204 examines the basic principles of protein structure, protein purification and characterization, proteomics, enzyme kinetics and mechanism, membrane structure and function, metabolism, and cellular energy transduction. The primary objective is to provide life scientists with an appreciation of basic principles of modern biochemistry, and of how the current conceptual and technical framework arose. Emphasis is placed on the experimental approaches and reasoning behind the dissection and reconstitution of these processes in a biological and, in some cases, clinical context. Discussions directed at biochemical problem solving, experimental design and the application of quantitative methods are integral to the course.
BIOL 3004-601	Infectious Disease Biology	Nataliya Balashova			This course focuses on selected topics concerning infectious agents, the diseases they cause in humans, and the social and scientific challenges they pose. The first section addresses the principles of epidemiology and microbial pathogenesis, as well as pathophysiology of infectious diseases. In the second section, tools and techniques of diagnosis, tracking, and control of infectious diseases will be discussed. To develop a broad understanding of the many different aspects of infectious processes, selected viral, fungal, protozoan, and helminthic pathogens and related infectious diseases will be presented. This course is open to students in the College of Liberal and Professional Studies only.
BIOL 3006-601	Histology	Brahim Chaqour		R 5:15 PM-8:14 PM	This course is designed to introduce the undergraduate student to the structure of tissues at the cellular level and to the way in which those tissues are assembled into organs. This knowledge of structure will be the basis for discussion of tissue and organ function. This course is open to students in the College of Liberal and Professional Studies only.
BIOL 3313-601	Essentials of Pathophysiology	Xiaohong Witmer		TR 5:15 PM-6:44 PM	This course is a study of homeostatic changes that occur with disease, and the implications of those changes in the progression and treatment of disease at molecular and cellular levels. Generalized mechanisms of disease as well as diseases of individual organ systems will be examined, with a view to understanding homeostatic compensations that occur as a result of altered function.
BIOL 3625-001	Marine Biology	Katie Lynn Barott Sherwood		TR 1:45 PM-3:14 PM	An introduction to marine biology and oceanography. Topics will include chemical and physical oceanography, a survey of form, function and phylogeny of algae, invertebrates and vertebrates, and an examination of ecological and evolutionary principles as applied to marine organisms and ecosystems.
BIOL 3711-401	Microbial Diversity and Pathogenesis Lab	Karl G Siegert		TR 12:00 PM-2:59 PM	The importance of microbiology in complex issues, such as the impact of the microbiome in human health or as alternative energy sources, is being appreciated more and more each day. This upper level laboratory course provides students with a robust technical skill set while also giving them an opportunity to participate in an authentic research project that may lead to novel discoveries. Students will generate research questions, formulate hypotheses, design experiments, analyze data, and present their research findings to the class. In each project, students will use the cutting edge approach of metagenomics to evaluate the microbial diversity of their environment via Next Generation Sequencing. Students will also examine the function of microbial species within their communities. Potential projects include the isolation of novel antibiotic producers and the antibiotic they produce, designing and optimizing microbial fuel cells that can be used to generate electricity, or isolating antibiotic resistant bacteria and attempting novel approaches to inhibit or prevent their growth.	BIOL5711401
BIOL 3851-301	Biology and Society (SNF Paideia Program Course)	Mechthild Pohlschroder Paul Robert Schmidt		T 3:30 PM-6:29 PM	This course uses a biological foundation to explore general issues at the interface of biology and society. We will use both historical and contemporary reading materials, with an emphasis on the primary scientific literature, to inform discussions on often controversial issues in biology as well as the social responsibility of scientists to respond to these issues. The course will cover how science has shaped social and political opinions on such topics as race, ethnicity, and gender, as well as how society and politics are influenced by and impact science. This course will provide a background and context in which to consider, anticipate, and respond to biology's present and future ethical and social implications.
BIOL 4004-601	Immunobiology	Michael P Cancro		M 7:00 PM-9:59 PM	Early development of microbiology, pathology, and immunobiology; molecular and cellular bases of immune phenomena including: immunity to pathogens, immune diseases, autoimmunity, and hypersensitivity. This course is open to students in the College of Liberal and Professional Studies only.
BIOL 4007-301	Cancer Cell Biology	Brian D. Keith		TR 1:45 PM-3:14 PM	This course will focus on the molecular mechanisms by which fundamental cellular processes are disrupted in the development of cancer.
BIOL 4010-401	Advanced Cell Biology	Wei Guo		TR 10:15 AM-11:44 AM	This course is designed for beginning graduate students and advanced undergraduates with a particular enthusiasm for cell biology. Biology 4010 does not attempt to cover all aspects of cell biology, and is therefore not appropriate for students seeking a lecture course which provides a comprehensive survey of the field. Rather, the primary objective of this course is to teach those students considering a career in the biomedical sciences how to read, discuss, and question original research papers effectively. Intensive classroom discussions focus on the experimental methods used, results obtained, interpretation of these results in the context of cell structure and function, and implications for further studies.	BIOL5010401, CAMB4800401
BIOL 4010-601	Advanced Cell Biology	Xiaohong Witmer		TR 7:00 PM-8:29 PM	This course is designed for beginning graduate students and advanced undergraduates with a particular enthusiasm for cell biology. Biology 4010 does not attempt to cover all aspects of cell biology, and is therefore not appropriate for students seeking a lecture course which provides a comprehensive survey of the field. Rather, the primary objective of this course is to teach those students considering a career in the biomedical sciences how to read, discuss, and question original research papers effectively. Intensive classroom discussions focus on the experimental methods used, results obtained, interpretation of these results in the context of cell structure and function, and implications for further studies.	CAMB4800601
BIOL 4016-401	Molecular Mechanisms of Infectious Disease Biology	David S. Roos			This course is designed for advanced undergraduates and beginning graduate students with a particular interest in infectious disease biology. Note that this course is not a comprehensive survey of the field and is not appropriate for students seeking a lecture course on disease. The primary objective of this course is to teach students considering a career in the biomedical sciences how to read, discuss, and question research papers effectively. Intensive classroom discussions focus on the experimental methods used, results obtained, interpretation of these results in the context of pathogen interactions with host cells and organisms, and implications for basic research and therapeutic development.	BIOL5016401
BIOL 4018-301	Cell Communication and Disease	John D Wagner		MW 3:30 PM-4:59 PM	Effective coordination between cells through cell communication and signaling enables multicellular organisms to develop and survive. Conversely, aberrations in these pathways are at the heart of a wide variety of human diseases. In this seminar course, we will discuss the molecular and cellular mechanisms of cell communication using a series of human diseases as a framework. The course will introduce postbac and advanced undergraduate students to the fundamental principles of cell signaling and will explore current questions of interest to the field. The synergistic nature of research directed at understanding basic cell biology, development and physiology with research aimed at elucidation and control of specific human ailments will be emphasized. The course will be comprised of a combination of introductory lectures and extensive discussion of primary literature. Students are expected to have a basic knowledge of cell biology, biochemistry and cell structure. BIOL 2201 and BIOL 2801 are recommended pre-requisites.
BIOL 4018-601	Cell Communication and Disease	John D Wagner		CANCELED	Effective coordination between cells through cell communication and signaling enables multicellular organisms to develop and survive. Conversely, aberrations in these pathways are at the heart of a wide variety of human diseases. In this seminar course, we will discuss the molecular and cellular mechanisms of cell communication using a series of human diseases as a framework. The course will introduce postbac and advanced undergraduate students to the fundamental principles of cell signaling and will explore current questions of interest to the field. The synergistic nature of research directed at understanding basic cell biology, development and physiology with research aimed at elucidation and control of specific human ailments will be emphasized. The course will be comprised of a combination of introductory lectures and extensive discussion of primary literature. Students are expected to have a basic knowledge of cell biology, biochemistry and cell structure. BIOL 2201 and BIOL 2801 are recommended pre-requisites.
BIOL 4026-401	Chromosomes and the Cell Cycle	Michael A. Lampson		TR 8:30 AM-9:59 AM	Life depends on the propagation of genetic material from one generation to the next through cycles of genome replication and cell division. The genome is copied by the parent, and one exact copy is inherited by each daughter cell. We will treat chromosomes as discrete entities, rather than collections of genes, that are replicated and divided with high fidelity to ensure that the genome remains stable over many generations. By reading selected primary literature covering several decades, we will build an understanding of the cell cycle by focusing on chromosomes and the associated molecular machinery. We will explore mechanisms that underlie replication and division, particularly control mechanisms that maintain genome integrity and are critical to prevent disease. The goal of the course is to develop a picture of the cell cycle by examining some of the key experiments and insights that have led to our current understanding.	BIOL5026401, CAMB4860401				https://coursesintouch.apps.upenn.edu/cpr/jsp/fast.do?webService=syll&t=202410&c=BIOL4026401
BIOL 4116-401	Neural Circuits for Survival	John Nicholas Betley		W 3:30 PM-6:29 PM	A fundamental goal of neuroscience is to understand how neural circuits in the brain function to influence behavior. The aim of this course is to highlight the neural basis of behavior and discuss modern approaches and novel methods to study the neuronal control of classically studied aspects of behavior. Through a combination of discussions, student presentations, and interactive lectures, we will explore the neural systems that regulate the interactions an animal has with the external world. We will explore sensory systems (such as vision, taste, and olfaction), motor systems, and survival behaviors (such as feeding, drinking, mating, and aggression). The course evaluation will be based largely on written work, participation, and presentations.	BIOL5116401
BIOL 4119-401	Biological Basis of Animal Diversity	Yun Ding		T 3:30 PM-6:29 PM	Animals display extraordinary diversity in their morphology, physiology, and behavior. Traditionally, these topics have been mostly studied from an ecological perspective. This course will focus on recent advances and discoveries that address the underlying biological mechanisms of animal diversity. Specific topics will include the genetic, molecular, and developmental basis of animal morphological diversity, and genetic, molecular, and neural basis of animal behavioral diversity. Students will gain an understanding of how animal diversity is encoded at the different levels of biological organization. The course will be comprised of lectures to introduce topics, discussion of primary literature, and in-class activities.	BIOL5119401
BIOL 4231-401	Genome Science and Genomic Medicine	Brian Gregory		TR 1:45 PM-3:14 PM	This course will be a focused study of genomes, genomic techniques, and how these approaches are and will be used in diagnosing and treating human disease. Topics will include genome sequencing, analysis of sequences and microarrays, and new techniques including high-throughput sequencing and reverse genetic analysis with a focus on genome-wide mutant collections.	BIOL5231401, CAMB4310401
BIOL 4233-401	Genes in Conflict: How sex and selfish genetic elements drive genome evolution	Mia Tauna Levine			In this course we explore the genetic basis of adaptation. We will investigate the forces that drive adaptation (e.g., environmental stress, pathogens, conflict, sex), the genetic mechanisms of adaptation (protein sequence changes, expression divergence, gene duplication, etc.), and the consequences of adaptation for contemporary cellular functions, developmental processes, and ecological interactions. The class meetings will be structured around both lectures and student-led discussions of the primary literature.	BIOL5233401
BIOL 4234-401	Epigenetics	Doris Wagner		MW 1:45 PM-3:14 PM	This course investigates epigenetic phenomena: heritable alternate states of gene activity that do not result from an alteration in nucleotide composition (mutations). Epigenetic mechanisms regulate genome accessibility and cell differentiation. They play a key role in normal development and in oncogenesis. For example both mammalian X-chromosome inactivation and nuclear transfer (cloning) are subject to epigenetic regulation. Amongst the epigenetic mechanisms we will discuss in this course are chromatin organization, histone modification, DNA methylation and non-coding RNAs. The course is geared toward advanced undergraduate and beginning graduate students and is a combination of lectures, student presentations and research presentations by guest speakers. Students will work with the current scientific literature.	BIOL5234401, CAMB4830401
BIOL 4250-401	Molecular Genetics of Development	Aman Yebio Husbands		M 10:15 AM-1:14 PM	Development is the process by which organisms grow and acquire their final shape. This remarkably complex process requires exquisite spatiotemporal control, and principles of developmental biology have implications for nearly all other biological disciplines. This course is a deep dive into these general biological principles, using plants as a model system. Students will prepare presentations on primary literature and engage in vigorous discussions in a "journal club" format. Our goal is to learn how developmentally significant genes and cellular interactions control differentiation and pattern formation.	BIOL5250401
BIOL 4318-401	Systems Biology: Integrative physiology and biomechanics of the muscular system	Lawrence Rome		TR 1:45 PM-3:14 PM	The course will focus on muscle function from the level of molecules to whole animal locomotion. At each level of organization, muscle function will be explored from mechanical and energetic viewpoints. The course will include lectures, demonstrations, and several guest expert lectures. Students will also be introduced to realistic musculo-skeletal modelling and forward dynamic simulations to explore integrated function.	BIOL5318401
BIOL 4517-401	Theoretical Population Biology	Erol Akcay			Introduction to basic theoretical tools to study the evolutionary and ecological dynamics of populations. Topics to be discussed include: basic population dynamics and population genetics theory, evolutionary game theory/adaptive dynamics, social evolution (kin selection/multilevel selection), life-history evolution, and stochastic models. Other topics may be added based on the specific interests of students in the class.	BIOL5517401
BIOL 4615-401	Freshwater Ecology	Melinda Daniels		MW 3:30 PM-4:59 PM	Survey of the physical, chemical and biological properties of freshwater ecosystems, both riverine and lentic, natural and polluted.	BIOL5615401, ENVS2390401
BIOL 4669-401	Plant Physiology Through Space and Time	Brent R. Helliker		F 12:00 PM-2:59 PM	This course is a lab/lecture/seminar hybrid that will meet once per week for three hours. Each session will consist of mini-lecture/lab, paper discussions/lab, or solely lab efforts. All reading assignments will be available on Canvas (no textbook fees). We will exam various aspects of photosynthesis, water relations and nutrient acquisition in the context of the evolutionary progression of higher plants. With each subject, we will consider, measure, and in some cases model whole-plant physiology while examining sub-cellular-level controls and ecosystem-to-global-level consequences. This course is designed to give molecular biologists through earth-system scientists the tools to measure and understand whole-plant physiological responses to molecular manipulation and environmental variability. All students will learn to appreciate the context of their work on both micro and macro scales.	BIOL5669401
BIOL 4710-401	Topics in Prokaryotic Biology: From Molecules to Microbiomes	Mark D Goulian		TR 8:30 AM-9:59 AM	This course will cover research articles from both the classic and contemporary literature on the genetics, cell biology, and physiology of prokaryotes. The material will focus on a small number of subjects in depth, with an emphasis on how the field has arrived at its current state of knowledge and on exciting new research directions. Possible topics include: stress responses, cell signaling, subcellular organization, bacteriophages, microbial communities, and host-microbe interactions.	BIOL5720401
BIOL 4825-401	Biochemistry and Molecular Genetics Superlab	John D Wagner		TR 1:45 PM-4:44 PM	Intensive laboratory class where open-ended, interesting biological problems are explored using modern lab techniques. Topics may include protein structure/function studies; genetic screens, genomics and gene expression studies; proteomics and protein purification techniques; and molecular cloning and DNA manipulation. The course emphasizes developing scientific communication and independent research skills. Course topics reflect the interests of individual Biology faculty members. This course is recommended for students considering independent research.	BIOL5825401
BIOL 5010-401	Advanced Cell Biology	Wei Guo		TR 10:15 AM-11:44 AM	This course is designed for beginning graduate students and advanced undergraduates with a particular enthusiasm for cell biology. Biology 4010/5010 does not attempt to cover all aspects of cell biology, and is therefore not appropriate for students seeking a lecture course which provides a comprehensive survey of the field. Rather, the primary objective of this course is to teach those students considering a career in the biomedical sciences how to read, discuss, and question original research papers effectively. Intensive classroom discussions focus on the experimental methods used, results obtained, interpretation of these results in the context of cell structure and function, and implications for further studies.	BIOL4010401, CAMB4800401
BIOL 5016-401	Molecular Mechanisms of Infectious Disease Biology	David S. Roos			This course is designed for advanced undergraduates and beginning graduate students with a particular interest in infectious disease biology. Note that this course is not a comprehensive survey of the field and is not appropriate for students seeking a lecture course on disease. The primary objective of this course is to teach students considering a career in the biomedical sciences how to read, discuss, and question research papers effectively. Intensive classroom discussions focus on the experimental methods used, results obtained, interpretation of these results in the context of pathogen interactions with host cells and organisms, and implications for basic research and therapeutic development.	BIOL4016401
BIOL 5026-401	Chromosomes and the Cell Cycle	Michael A. Lampson		TR 8:30 AM-9:59 AM	Life depends on the propagation of genetic material from one generation to the next through cycles of genome replication and cell division. The genome is copied by the parent, and one exact copy is inherited by each daughter cell. We will treat chromosomes as discrete entities, rather than collections of genes, that are replicated and divided with high fidelity to ensure that the genome remains stable over many generations. By reading selected primary literature covering several decades, we will build an understanding of the cell cycle by focusing on chromosomes and the associated molecular machinery. We will explore mechanisms that underlie replication and division, particularly control mechanisms that maintain genome integrity and are critical to prevent disease. The goal of the course is to develop a picture of the cell cycle by examining some of the key experiments and insights that have led to our current understanding.	BIOL4026401, CAMB4860401				https://coursesintouch.apps.upenn.edu/cpr/jsp/fast.do?webService=syll&t=202410&c=BIOL5026401
BIOL 5116-401	Neural Circuits for Survival	John Nicholas Betley		W 3:30 PM-6:29 PM	A fundamental goal of neuroscience is to understand how neural circuits in the brain function to influence behavior. The aim of this course is to highlight the neural basis of behavior and discuss modern approaches and novel methods to study the neuronal control of classically studied aspects of behavior. Through a combination of discussions, student presentations, and interactive lectures, we will explore the neural systems that regulate the interactions an animal has with the external world. We will explore sensory systems (such as vision, taste, and olfaction), motor systems, and survival behaviors (such as feeding, drinking, mating, and aggression). The course evaluation will be based largely on written work, participation, and presentations.	BIOL4116401
BIOL 5119-401	Biological Basis of Animal Diversity	Yun Ding		T 3:30 PM-6:29 PM	Animals display extraordinary diversity in their morphology, physiology, and behavior. Traditionally, these topics have been mostly studied from an ecological perspective. This course will focus on recent advances and discoveries that address the underlying biological mechanisms of animal diversity. Specific topics will include the genetic, molecular, and developmental basis of animal morphological diversity, and genetic, molecular, and neural basis of animal behavioral diversity. Students will gain an understanding of how animal diversity is encoded at the different levels of biological organization. The course will be comprised of lectures to introduce topics, discussion of primary literature, and in-class activities.	BIOL4119401
BIOL 5210-401	Molecular Biology and Genetics	Aman Yebio Husbands Mia Tauna Levine		TR 10:15 AM-11:44 AM	This course will survey the discipline of molecular genetics. Two broad areas will be considered 1) Molecular Biology: DNA replication, transcription, translation, regulation of gene expression in both prokaryotic and eukaryotic systems, and genomics and 2) Genetics: basic Mendelian & molecular genetics.	BIOL2210401
BIOL 5210-402	Molecular Biology and Genetics			W 12:00 PM-1:29 PM	This course will survey the discipline of molecular genetics. Two broad areas will be considered 1) Molecular Biology: DNA replication, transcription, translation, regulation of gene expression in both prokaryotic and eukaryotic systems, and genomics and 2) Genetics: basic Mendelian & molecular genetics.	BIOL2210402
BIOL 5210-403	Molecular Biology and Genetics			W 5:15 PM-6:44 PM	This course will survey the discipline of molecular genetics. Two broad areas will be considered 1) Molecular Biology: DNA replication, transcription, translation, regulation of gene expression in both prokaryotic and eukaryotic systems, and genomics and 2) Genetics: basic Mendelian & molecular genetics.	BIOL2210403
BIOL 5220-401	Human Evolutionary Genomics	Sarah Anne Tishkoff			Advanced seminar on current topics in human genomics and human evolution. Topics include the methods used for mapping and sequencing genomes; phylogenetic and population genetic analysis; and detecting variation in the human genome. This course is designed for graduate students but advanced undergraduates with a strong background in genetics are also welcome.	CAMB5220401
BIOL 5231-401	Genome Science and Genomic Medicine	Brian Gregory		TR 1:45 PM-3:14 PM	This course will be a focused study of genomes, genomic techniques, and how these approaches are and will be used in diagnosing and treating human disease. Topics will include genome sequencing, analysis of sequences and microarrays, and new techniques including high-throughput sequencing and reverse genetic analysis with a focus on genome-wide mutant collections.	BIOL4231401, CAMB4310401
BIOL 5233-401	Genes in Conflict: How sex and selfish genetic elements drive genome evolution	Mia Tauna Levine			In this course we explore the genetic basis of adaptation. We will investigate the forces that drive adaptation (e.g., environmental stress, pathogens, conflict, sex), the genetic mechanisms of adaptation (protein sequence changes, expression divergence, gene duplication, etc.), and the consequences of adaptation for contemporary cellular functions, developmental processes, and ecological interactions. The class meetings will be structured around both lectures and student-led discussions of the primary literature.	BIOL4233401
BIOL 5234-401	Epigenetics	Doris Wagner		MW 1:45 PM-3:14 PM	This course investigates epigenetic phenomena: heritable alternate states of gene activity that do not result from an alteration in nucleotide composition (mutations). Epigenetic mechanisms regulate genome accessibility and cell differentiation. They play a key role in normal development and in oncogenesis. For example both mammalian X-chromosome inactivation and nuclear transfer (cloning) are subject to epigenetic regulation. Amongst the epigenetic mechanisms we will discuss in this course are chromatin organization, histone modification, DNA methylation and non-coding RNAs. The course is geared toward advanced undergraduate and beginning graduate students and is a combination of lectures, student presentations and research presentations by guest speakers. Students will work with the current scientific literature.	BIOL4234401, CAMB4830401
BIOL 5250-401	Molecular Genetics of Development	Aman Yebio Husbands		M 10:15 AM-1:14 PM	Development is the process by which organisms grow and acquire their final shape. This remarkably complex process requires exquisite spatiotemporal control, and principles of developmental biology have implications for nearly all other biological disciplines. This course is a deep dive into these general biological principles, using plants as a model system. Students will prepare presentations on primary literature and engage in vigorous discussions in a "journal club" format. Our goal is to learn how developmentally significant genes and cellular interactions control differentiation and pattern formation.	BIOL4250401
BIOL 5318-401	Systems Biology: Integrative physiology and biomechanics of the muscular system	Lawrence Rome		TR 1:45 PM-3:14 PM	The course will focus on muscle function from the level of molecules to whole animal locomotion. At each level of organization, muscle function will be explored from mechanical and energetic viewpoints. The course will include lectures, demonstrations, and several guest expert lectures. Students will also be introduced to realistic musculo-skeletal modelling and forward dynamic simulations to explore integrated function.	BIOL4318401
BIOL 5410-401	Evolutionary Biology	Paul Robert Schmidt Corlett Wood		TR 10:15 AM-11:44 AM	Theories and mechanisms of evolution, with emphasis on the genetic basis of evolutionary change.	BIOL2410401
BIOL 5410-406	Evolutionary Biology			R 12:00 PM-12:59 PM	Theories and mechanisms of evolution, with emphasis on the genetic basis of evolutionary change.	BIOL2410406
BIOL 5517-401	Theoretical Population Biology	Erol Akcay			Introduction to basic theoretical tools to study the evolutionary and ecological dynamics of populations. Topics to be discussed include: basic population dynamics and population genetics theory, evolutionary game theory/adaptive dynamics, social evolution (kin selection/multilevel selection), life-history evolution, and stochastic models. Other topics may be added based on the specific interests of students in the class.	BIOL4517401
BIOL 5615-401	Freshwater Ecology	Melinda Daniels		MW 3:30 PM-4:59 PM	Survey of the physical, chemical and biological properties of freshwater ecosystems, both riverine and lentic, natural and polluted.	BIOL4615401, ENVS2390401
BIOL 5669-401	Plant Physiology Through Space and Time	Brent R. Helliker		F 12:00 PM-2:59 PM	This course is a lab/lecture/seminar hybrid that will meet once per week for three hours. Each session will consist of mini-lecture/lab, paper discussions/lab, or solely lab efforts. All reading assignments will be available on Canvas (no textbook fees). We will exam various aspects of photosynthesis, water relations and nutrient acquisition in the context of the evolutionary progression of higher plants. With each subject, we will consider, measure, and in some cases model whole-plant physiology while examining sub-cellular-level controls and ecosystem-to-global-level consequences. This course is designed to give molecular biologists through earth-system scientists the tools to measure and understand whole-plant physiological responses to molecular manipulation and environmental variability. All students will learn to appreciate the context of their work on both micro and macro scales.	BIOL4669401
BIOL 5711-401	Microbial Diversity and Pathogenesis Lab	Karl G Siegert		TR 12:00 PM-2:59 PM	The importance of microbiology in complex issues, such as the impact of the microbiome in human health or as alternative energy sources, is being appreciated more and more each day. This upper level laboratory course provides students with a robust technical skill set while also giving them an opportunity to participate in an authentic research project that may lead to novel discoveries. Students will generate research questions, formulate hypotheses, design experiments, analyze data, and present their research findings to the class. In each project, students will use the cutting edge approach of metagenomics to evaluate the microbial diversity of their environment via Next Generation Sequencing. Students will also examine the function of microbial species within their communities. Potential projects include the isolation of novel antibiotic producers and the antibiotic they produce, designing and optimizing microbial fuel cells that can be used to generate electricity, or isolating antibiotic resistant bacteria and attempting novel approaches to inhibit or prevent their growth.	BIOL3711401
BIOL 5720-401	Topics in Prokaryotic Biology: From Molecules to Microbiomes	Mark D Goulian		TR 8:30 AM-9:59 AM	This course will cover research articles from both the classic and contemporary literature on the genetics, cell biology, and physiology of prokaryotes. The material will focus on a small number of subjects in depth, with an emphasis on how the field has arrived at its current state of knowledge and on exciting new research directions. Possible topics include: stress responses, cell signaling, subcellular organization, bacteriophages, microbial communities, and host-microbe interactions.	BIOL4710401
BIOL 5825-401	Biochemistry and Molecular Genetics Superlab	John D Wagner		TR 1:45 PM-4:44 PM	Intensive laboratory class where open-ended, interesting biological problems are explored using modern lab techniques. Topics may include protein structure/function studies; genetic screens, genomics and gene expression studies; proteomics and protein purification techniques; and molecular cloning and DNA manipulation. The course emphasizes developing scientific communication and independent research skills. Course topics reflect the interests of individual Biology faculty members. This course is recommended for students considering independent research.	BIOL4825401
BIOL 6010-301	Communication for Biologists	Brian Gregory Joshua Benjamin Plotkin			Basic science writing and presentation skills for PhD students in Biology. Designed for second year graduate students preparing for qualifying exams. In the first half of the course, students will produce weekly writing assignments and critique writing submitted by others. In the second half, students will learn techniques for effective research presentations in both seminar style environments and chalk-talk settings.